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332 Cards in this Set

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What are the four traits that Charophytes (plant's closest relative) and Plants have in common which distinguish them from other things (algae...etc.)?
1) Rosette-shaped cellulose-synthesizing complexes
2) Peroxisome enzymes
3) Flagellated sperm
4) Formation of phragmoplast
Who are plant's closest relatives?
Charophytes
What distinguishes charophytes (plant's closest relative) from plants?
Plants are terrestrial.
What are the four derived characteristics of plants?
1) Alternation of generations
2) Multicellular gametangia
3) Apical meristems
4) Sporopollenin
What is the significance of the "phragmoplast"?
Shared characteristic of charophytes and plants.
A group of microtubules that forms between daughter nuclei of a dividing cell which directs the cell plate formation.
What is the function of "peroxisome enzymes"?
Help minimize the loss of organic products as a result of photorespiration.
What are the two generations that alternate in plants?
Gametophyte (n)
Sporophyte (2n)
Starting with the gametophyte, what is the general life cycle of plants? (5 steps)
1) Gametophyte produces haploid gametes by mitosis
2) Two gametes unite (fertilization) and form a diploid zygote (2n)
3) Zygote develops into multicellular diploid sporophyte (2n)
4) Sporophyte produces haploid spores (n) by meiosis
5) Spores develop into multicellular haploid gametophytes (n) through mitosis
What are some advantages to being aquatic (charophytes)?
- Abundance of water
- Water is a buffer
- Structural support
- Reproduction is easy
What are some disadvantages to being aquatic (charophytes)?
- Dilution of nutrients
- Light is not as available
- Limited O2 and CO2
- No control of movement
What are some advantages to being terrestrial (plants)?
- More light available
- Minerals are more abundant (not diluted)
- O2 and CO2 are not limiting
- Stability in ground / anchor
- No competition initially
What are some disadvantages to being terrestrial (plants)?
- Extreme environments
- H2O is limiting
- Reproduction is more difficult
- Herbivores
- Lack of structural support
- UV radiation damage
Which are the nonvascular plants?
Bryophytes (liverworts, hornworts, mosses)
What can vascular plants be further divided into?
Seedless (lycophytes and pterophytes)
With seeds (gymnosperms and angiosperms)
What kind of seedless vascular plants are there?
Lycophytes (club mosses, spike mosses, quillworts)
Pterophytes (ferns, horsetails...)
What is the function of sporopollenin in plants?
Makes walls of plant spores tough and resistant to harsh environments; protects from desiccation (drying out).
What is the function of the cuticle?
Waxy, protective layer that prevents excess water loss from epidermis.
What is the function of lignin?
Provides structural support in terrestrial species.
What distinguishes bryophytes from other land plants?
No vascular system; most similar to ancestral plants; gametophyte is dominant (not sporophyte).
What are apical meristems?
Localized regions of cell division at the tips of shoots and roots that are undifferentiated.
What is the function of vascular tissue?
Cells that are joined into tubes that transport water and nutrients through the plant body.
What is the intermediary step between bryophyte spores and their dominant phase (gametophyte)?
Protonemata (n) - a mass of green branched, one cell thick filaments that grow from spores and buds to produce gametophytes.
Bryophyte (moss) gametophytes can be male or female... What are they called? How do they fertilize?
Male - Antheridia
Female - Archegonia
Sperm released from antheridia swim through a film of moisture to reach the egg.
Since bryophytes do not have roots, what do they rely on to anchor the gametophyte?
Rhizoids
What do the seta and capsule make up in a bryophyte (moss)?
Sporophyte
Why do mosses / bryophytes require a moist habitat?
Fertilization requires a film of water in order for sperm to swim to egg.
At the end of the sporophyte capsule in bryophytes / mosses, there is what structure which allows for the slow release of spores?
Peristome
What is the function of stomata?
Specialized pores which allow the exchange of CO2 and O2; also the route through which water evaporates from the sporophyte.
What is the origin of "peat"? What can it be used for?
Partially decayed organic material; fuel source (400 billion tons of C is sequestered).
How much Carbon do humans release per year?
7-8 billion tons per year.
What structures do vascular plants specifically have?
Xylem, Phloem, Leaves, Roots
What is the dominant generation in vascular plants?
Sporophyte
What is the difference between the terms homosporous and heterosporous?
Homosporous = one type of sporangium = one spore --> bisexual gametophyte
Heterosporous = two types of sporangia = differentiation of spores (megaspores, f, and microspores, m)
What are the five stages in the life cycle for a fern (seedless vascular plant)?
1) Sporangia release spores (homosporous)
2) Gametophyte produces eggs or sperm
3) Sperm use flagella to swim from the antheridia to eggs in the archegonia for fertilization
4) Zygote develops into a new sporophyte
5) On underside of sporophyte's reproductive leaves are spots called sori made up of a cluster of sporangia.
Where are the sporangium (spore producing organs) located on a fern?
On the underside of the sporophyte's reproductive leaves in spots called sori (each sorus is a cluster of sporangia).
Why did the gametophyte evolve to be less dominant and a smaller structure that is enclosed?
To protect the gametes from UV radiation and other harm.
The club mosses, spike mosses, and quillworts are what kind of plants?
Lycophytes: seedless vascular plants
The ferns, whisk ferns and horsetails represent which kind of plants?
Pterophytes (seedless vascular plants)
The ratio of CO2 in the atmosphere 600-400 mya was very high, what explained the crashing to today's level around 350 mya?
CO2 was being incorporated into plants
- evolution of vascular tissue, roots, and leaves increased the rate of photosynthesis
- coal was made by build up of carbon in plants under lots of pressure
- during Carboniferous period
When the rate of CO2 dropped due to increased photosynthesis, what also occurred in the atmosphere? What effect did this have?
O2 increased in atmosphere
Led to huge insects
What is a seed? What did seeds replace?
An embryo packaged with a supply of nutrients inside a protective coat; spores.
What are the two kinds of seed plants? How are they distinguished from one another?
Gymnosperms = naked seed plants
Angiosperms = ovaries / flowers and fruits
What three components make up the ovule?
Megosporangium, megaspore, integument
What is the male gametophyte for seed plants?
Pollen grain.
What forms the hard seed coat?
The integument becomes the seed coat.
What polymer protects the pollen grain / male gametophyte?
Sporopollenin.
Gymnosperms show the evolutionary transition to non-motile spores. How do they compensate for this?
The pollen grains are able to travel by the wind or animals, eliminating the need to swim via water.
What is the purpose of the pollen tube?
It discharges the sperm from the pollen grain into the egg.
What is the opening in the integument through which the pollen grain enters to fertilize the egg?
Micropyle
What are five characteristics of seed plants?
1) Reduced gametophytes
2) Heterosporous (two kinds of sporangia/spores)
3) Ovules (containing female gametophyte)
4) Pollen (containing male gametophyte)
5) Seeds
4)
Why are seeds more advantageous than spores?
- Seeds are more protected than spores (higher survival)
- Independent and dispensable from parent
What are the three components of a seed? From what do they originate from? Haploid or diploid?
- Integument (protective coat from parent sporophyte) (2n)
- Food supply (from parent gametophyte) (n)
- Embryo (new sporophyte) (2n)
What are the four kinds of gymnosperms?
1) Conifers (biggest phylum)
2) Cycads (2nd biggest phylum)
3) Ginkgo
4) Gnetophytes
What was the transitional species between seedless plants and seed plants that was heterosporous, no seeds, very developed vascular system
Progymnosperm (380 mya)
What are the four main structural parts of a flowering plant?
1) Sepal
2) Petal
3) Stamen (Anther & Filament)
4) Carpel (Stigma, Style, & Ovary)
What is a fruit?
Mature ovary.
What does the stamen develop into in angiosperms?
Pollen grain.
What does the carpel develop into in angiosperms?
Female gametophytes (embryo sac)
Which part of the flower receives the pollen? Which part connects the stigma to the ovary? Which part contains the ovules?
The stigma of the carpel. Style. Ovary.
What is meant by double fertilization?
One sperm fertilizes the egg to form the zygote.
One sperm produces a triploid cell (2 nuclei and 1 sperm)
What distinguishes Monocots from Dicots?
Monocots (vs. Dicots):
- One cotyledon (vs. two)
- Parallel veins (vs. netlike)
- Vascular tissue scattered in stem (vs. in rings)
- Root system is fibrous, no main root (vs. taproot, main root)
- Pollen grain has one opening (vs. three openings)
- Floral organs in multiples of 3 (vs. 4 or 5)
Spores are produced after which step? What do they produce?
After meiosis; give rise to gametophytes.
What are the three general components/organs of plants?
Roots (root system)
Stems & Leaves (shoot system)
Roots have evolved in almost all _____ plants.
Vascular
What is the purpose of roots?
Acquire water and minerals via root hairs; anchor plants; store carbohydrates.
What are some modified roots?
Prop roots, Storage roots, Fibrous roots, Aerial roots, Pneumophores (obtain O2 in wet settings by sticking out of H2O), Buttress roots
Single, elongated cells that absorb water and minerals by increasing surface area, are called what?
Root Hairs
Where does primary growth occur?
Apical meristems - localized regions of cell growth at tips of shoots and roots.
What are the two modified stems? How are they distinguished?
Rhizomes = below ground stems
Stolons = above ground stems
What is "vegetative growth"?
Production of non-reproductive leaves, stems and roots.
What is the function of leaves?
Increase surface area of plant which aids in photosynthesis.
What are the three kinds of leaf growth?
Simple leaf (single undivided blade)
Compound leaf (blade of multiple leaflets)
Double Compound leaf (each leaflet is divided into smaller leaflets)
What are the hairlike outgrowths of the shoot epidermis that reduce water loss and reflect excess light; also provide defense against insects with toxic chemicals?
Trichomes
What kind of modified leaves are there?
- Tendrils (support for pea plants)
- Spines (cacti)
- Storage leaves (water)
- Reproductive leaves (grow and fall off to form new plants)
- Bracts (poinsettia)
What are the three layers of tissue?
1) Dermal tissue - outer protective covering
2) Ground tissue - inside of plant
3) Vascular tissue - transport minerals / water / sugars (xylem & phloem)
What tissue replaces the epidermis in woody plants?
Periderm
What is the cell wall made of? What is its purpose?
Made of cellulose; provides rigidity and strength.
What is the space between cell walls which contains pectin and pectic acid?
Middle lemella
What are the five types of plant cells?
1) Parenchyma cells
2) Collenchyma cells
3) Sclerenchyma cells
4) Xylem
5) Phloem
Which kind of plant cells have a large central vacuole, is engaged in many metabolic functions, and has the ability to differentiate into other cells?
Parenchyma cells
Which kind of plant cells have a thicker primary cell wall that varies in thickness giving the plant flexibility and stability? Plant it's in?
Collenchyma cells; celery
Which kind of plant cells are used for support due to their unusually thick cell walls; they are dead upon maturing...?
Sclerenchyma cells
What are the two kinds of sclerenchyma cells? What are they used for?
Schlereids (hardness of nutshells, pears)
Fibers (in linens)
Which kind of plant cells are used for movement of water and minerals?
Xylem
What two kinds of cells make up the xylem?
Tracheids (narrower pipes to move water from roots to rest of plant)
Vessels (wider with perforation plate on one end)
Through which structure do the tracheids and vessels of the xylem pass water horizontally?
Pits (areas where there is only primary cell wall)
Which plant cells are dead at functional maturity?
Sclerenchyma cells and Xylem
Which type of plant cells are responsible for the movement of organic nutrients (sugars, amino acids, etc.)?
Phloem
What two kinds of cells are in the phloem?
Sieve cells (facilitate flow of liquid from cell to cell)
Companion cells (non-conducting)
What structure allows the movement of sugars between phloem cells?
Plasmodesmata (connected by membrane)
What are the areas in plants that are perpetually embryonic tissues? What two kinds are there?
Meristems: apical (at top of plant, produces primary growth) or lateral (along stem, secondary growth)
What two structures make up the lateral meristems (secondary growth)?
Vascular cambium
Cork cambium
What is the function of the vascular cambium?
Adds layers of secondary xylem and phloem.
What is the function of the cork cambium?
Replaces epidermis with periderm.
What is the stuff external to the vascular system (stem/trunk) called?
Cortex
What is the structure that includes the vascular tissue (xylem and phloem) within the endodermis of a root or stem? (Circular core of roots)
Stele
Which of the vascular tissues grows inwards? Which grows outward? (from vascular cambium)
Inward = xylem
Outward = phloem
What is the internal core inside of the xylem called?
Pith
What makes up the bark?
Secondary Phloem and Periderm
What are the two kinds of mesophyll (ground tissue) cells in a leaf?
Palisade cells
Spongy cells
Why do the spongy cells have so many openings between them?
For gas exchange (take up CO2, release O2)
Which cells focus the light on the chloroplasts?
Palisade cells
What's the difference between microphylls and megaphylls?
Microphyll = small, spine-shaped leaves with single strand of vascular tissue
Megaphyll = highly branched vascular systems (greater photosynthetic productivity)
When did plants first appear? Plants with leaves?
Plants = 475 mya
Leaves = 380 mya (coincided with drop in CO2 concentration)
In which tissue are chloroplasts located?
Mesophyll - tissue inside of leaf
Where does photosynthesis occur?
In chloroplasts in stacks of thylakoids in grana.
In the light reactions what are the inputs and outputs?
Inputs = H2O, light, NADP+, ADP
Outputs = O2, ATP, NADPH
In the dark reactions (Calvin Cycle) what are the inputs and outputs?
Inputs = CO2, ATP, NADPH, RuBP
Outputs = Sugars, NADP+, ADP
Where do the light reactions occur? Dark reactions?
Light = thylakoids
Dark = stroma (dense fluid in chloroplasts surrounding the thylakoid membranes)
What do the light reactions do?
Converts solar energy to chemical energy (ATP).
What do the dark reactions (Calvin Cycle) do?
Produces sugars only with help from NADPH and ATP that are generated during the light reactions.
What is the most abundant protein? What is it's job?
Rubisco - fixes CO2
What are the three basic steps of the Calvin Cycle?
1) Carbon fixation (3 CO2 + Rubisco --> 3-phosphoglycerate)
2) Reduction (form G3P, one released as sugar)
3) Regeneration of CO2 acceptor (RuBP via 5 G3P)
When does photorespiration occur?
As the concentration of O2 increases the rate of photosynthesis decreases and photorespiration increases.
What molecule is responsible for photorespiration occurring? Why?
Rubisco is reactive with O2 as well as CO2; evolved back when O2 had a low concentration, so it was not necessary to be exclusive of O2
What happens when photorespiration is occurring?
Consumes ATP, produces no sugars, decreases photosynthetic output.
Which enzyme can be utilized instead of rubisco and is useful due to its ability to preferentially use CO2 and not O2?
PEP Carboxylase (adds PEP to CO2 to make oxaloacetate)
When PEP Carboxylase adds PEP to CO2 what forms?
Oxaloacetate --> Malate --> ...
The malate produced in the mesophyll cells (by PEP --> Oxaloacetate) is sent where to do what?
Bundle-Sheath Cells to enter Calvin Cycle (brings CO2)
The sugars made by the Calvin Cycle are released to where?
Vascular tissue / veins (for transport)
What distinguishes a C3 plant?
Stomates close on hot, dry days to conserve water; decreases the photosynthetic potential.
What distinguishes a C4 plant?
Alternate mode of carbon fixation; bundle sheath and mesophyll cells help to concentrate CO2 in Calvin Cycle; increases productivity in hot, dry areas.
In which kind of plants is the alternative PEP Carboxylase method utilized?
CAM and C4 plants (helps them work in hot, dry areas)
What distinguishes CAM plants?
- Alternate mode of carbon fixation
- Open stomates at night (C4 Cycle / CO2 fixation)
- Close stomates during day and close out O2 (Calvin Cycle / Sugar production)
Where are CAM plants usually found?
Harsh environments (hot / dry conditions) (deserts, succulent plants)
Why do C4 plants have an advantage in hot, dry conditions?
- No (or little) photorespiration
- PEP carboxylase scavenges CO2 at low concentrations so stoma closure has little effect
- Calvin Cycle runs to use all CO2
- Increased nitrogen efficiency
When do C3 plants have an advantage over C4 plants?
Lower leaf temperatures = higher net rate of photosynthesis
(as it gets warmer, C3 plants close stomates, slowing photosynthesis)
When do C4 plants have an advantage over C3 plants?
Higher leaf temperature, higher light intensities.
Where are most C4 plants found?
Tropical regions
What does primary growth produce?
Epidermis, ground tissue, and vascular tissue
In the roots, how can the stele be described?
Vascular cylinder
Where does secondary growth arise from?
Lateral meristems (vascular cambium and cork cambium)
What is the nonfunctional xylem wood in the center of the trunk called?
Heartwood
What is the layer around the heartwood called?
Sapwood
What layers make up the secondary xylem?
Hearwood, Sapwood (from vascular cambium)
How do plant cells divide?
Mitotically by putting down a cell plate
What is the function of vacuoles?
Take in water and hold it; expands cell
Why do plants have to asymmetrically divide?
So they can determine polarity / directionality
What determines where the cell plate goes?
Direction of the bands of microtubules around the nuclei (microfibrils align themselves according to direction of microtubules which determines which way the cell expands).
In which kind of mutant is the orientation of microtubules random (thus generating cells growing in all directions)?
Fass mutants
In which kind of mutant is the establishment of polarity defective?
Gnom
What are homeotic genes? What are the two we studied?
Genes that regulate development
- KNOTTED-1
- GLABRA-2
What is the homeotic gene which is involved in leaf morphology?
Knotted-1
What is the homeotic gene which is involved in root hair distribution?
Glabra-2
From where do plants acquire water and minerals?
Roots
From where do plants acquire energy/light and CO2?
Leaves
What is the main factor influencing the level of transpiration in a plant?
Water availability
What is the LAI? Value for a young plant? Common values for mature plants?
Leaf Area Index = ratio of area of leaves per area of ground
- Young: <1
- Mature: 5-7
What would be likely to happen if the Leaf Area Index (LAI) value was greater than 7?
More respiration would be occurring than photosynthesis; plant would self-prune to decrease the LAI.
What are mycorrhizae? What do mycorrhizae do for a plant?
Fungi that has a symbiotic relationship with plant which helps plants acquire water and nutrients in the roots.
What generates a voltage gradient for active transport in plants?
Proton pump
How does the cell move H+ from the cytoplasm out of the cell to the extracellular fluid?
ATP Hydrolysis which generates an EMF
Why do cells move H+ to the extracellular fluid?
Allows the cell to transport other sugars/ions/things into the cell via the EMF and transport proteins.
In what way does water prefer to flow?
From High-water potential to Low-water potential (low solute --> high solute)
What makes up the "water potential"?
Solute concentration and physical pressure (of cell wall, etc).
What is the term for the movement of water across a membrane?
Osmosis
How can the water potential be lowered?
Addition of solutes
What is the water potential equation?
ψ (w) = ψ (s) + ψ (p)
water potential = solute potential (always negative) + pressure potential (pos. or neg.)
When the pressure potential is 0, what does the cell look like? Why?
Flaccid - membrane is not pushed up to the cell wall
What is the solute potential proportional to?
Molarity (0.1M = -0.23 MPa)
What is happening in a plasmolyzed cell?
Water is leaving the cell and the membrane is trying to come in to try to stop the water loss.
What is the normal state of a plant cell?
Turgid - water is trying to enter the cell, cell wall is up against the membrane, cell wall i strong and rigid
What is loss of turgidity called in a plant cell?
Wilting
Pure water has a water potential of what? When there is a solute in a solution, how does this affect the solute potential?
Pure water = 0 MPa
Solute solution = negative MPa
How do plants move from cell to cell without crossing the membranes?
Via plasmadesmata (a continuous membrane connection with cytosol)
What is the symplastic route of movement between plant cells?
Crossing between cells connected by plasmadesmata.
What is the route that crosses cell membranes and walls when moving from cell to cell?
Apoplastic route
Which route goes through the symplast and apoplast?
Transmembrane route
What is the name of the barrier in the cell wall that prevents movement of water and nutrients between cells?
Casparian Strip
How do nutrients and water enter the stele if there is a Casparian Strip preventing their movement?
Must cross a selectively permeable barrier by entering the internal symplastic route.
What are the proteins that facilitate water movement? What is their purpose?
Aquaporins - speed up the rate of water crossing the phospholipid bilayer
What is bulk flow?
Long-distance movement of water driven by pressure, which is much faster than diffusion or active transport.
What can explain upward transport of water in a plant up to one meter high?
Root Pressure
What are the openings on leaves (modified pores) that allows water to escape by guttation due to root pressure?
Hydathodes
What is the term for water leaving the tips of leaves via the hydathodes (due to root pressure)?
Guttation
What explains water movement in a plant higher than 1m?
Water potential (high to low)
(gradient from high to low, roots to leaves)
What does cohesion refer to in regards to water movement?
Water molecules stick together in a column of water.
What does adhesion refer to in regards to water movement?
Water molecules stick to the cell wall to fight against gravity.
What state is water in within plants?
Liquid in plant, turns to gas when it reaches the stoma to evaporate.
What is cavitation? What does it result in?
Cavitation is the act of forming an embolism, an air pocket, which prevents the movement of water.
How do plants bypass embolisms (air pockets which prevent water movement)?
Via pits (openings in the cell wall) which allows water to bypass blockages.
When guard cells accumulate K+, what happens?
Stomata open
What is the structure of a stoma (plural stomata)?
Two guard cells
What four cues influences stomates opening/closing?
1) Light (triggers opening)
2) CO2 depletion
3) Internal clock (usually open during day)
4) ABA hormone signals from roots to tell stomates to close when water levels are low
When the stoma is open, guard cells are _____.
When the stoma is closed, guard cells are _____.
Open = turgid
Closed = flaccid
Organic compounds are produced in ____ and are transported to ____.
Sources
Sinks
How do organic compounds get transported?
Active transport: H+ pump (produces gradient) and cotransport (of sugar molecules with H+)
What are the four steps of the pressure-flow hypothesis?
1) Loading of sugar from source cell (leaf) into phloem (sieve tube)
2) Uptake of water into sieve tube; attempts to dilute
3) Unloading of sugar into sink cell (storage root)
4) H20 recycled (into xylem)
How do phloem and xylem differ in the direction of transport?
Xylem = unidirectional (from roots to leaves)
Phloem = bidirectional
What proved that sugar concentration is higher in sieve tubes closer to sugar sources?
Aphids which feed via stylets (stick into phloem and feed on sugar); helpful for measuring concentration of sucrose.
Why can xylem transport water and minerals with dead cells but phloem requires living cells?
-Water is moved along a gradient of water potential by tension in xylem (does not need to be alive).
-Phloem sap requires active movement of sugar to concentrate it
What do viral movement proteins influence?
Size of the plasmadesmata (want it to be larger to allow viral RNA to pass between cells)
Where do plant viruses move?
In phloem (prefer live cells to dead xylem cells)
What four components make up the composition of soil?
- Sand (large)
- Silt (small)
- Clay (very small)
- Humus (organic matter)
Which component of soil holds the least amount of water?
Sand (very porous)
Although clay can hold a lot of water, why might this be a bad thing?
Does not allow for flow of water.
What is the best composition of soil texture?
1/3 of each (sand, silt, clay) approximately
What is humus?
Remains of dead organisms and other organic matter which is resistant to decomposition; stable and long-lived.
What are the layers of soil called?
Horizons (A, B, C)
What is topsoil? Where is it fond?
A mixture of broken-down rock, living organisms, and humus; Horizon A.
(Water and wind erosion remove considerable amounts)
What is the best kind of soil?
Loam (most fertile, has a roughly equivalent amount of each)
What charge do soil particles have? Why is this significant?
Negative charged at surface; attract positive ions (cations) like K+, Cu2+, Mg2+, Ca2+, H+
What is the CEC?
Cation Exchange Capacity - different soils have differing abilities to attract/give up these cations depending on the number of cation adhesion sites and pH.
How does cation exchange occur?
Root hairs release H+ from H2O; H+ ions are attracted to negative soil particles, bumping off mineral cations which are then absorbed by root hairs.
Which kind of ions are most susceptible to leeching?
Anions because cations are attracted to soil particles; anions sink through horizons and bed rock to the ground water when there is a lot of water in the soil.
Why must sand be irrigated way more often than a good soil like silt loam?
Sand can hold 0.7 inches of water
Silt loam can hold 2.5 inches of water
(On hot days, can lose up to 0.3 inches)
What is tillage used for?
Controlling weeds
What are negative consequences of tilling?
Disrupts soil (roots are disturbed that hold soil in place leading to increased runoff and erosion)
Compaction (soil particles are pressed together, reducing pore space between them, reduces water and nutrient absorption and slows gas exchange and root growth)
What kind of experiments are used to determine what minerals are needed for plants?
Hydroponic growth (grow in water containing various minerals/nutrients, no soil)
What are the macro-nutrients plants require?
C, H, O, N, K, Ca, Mg, P, S
What are the micro-nutrients plants require?
Cl, Fe, Mn, B, Zn, Cu, Ni, MoO4
Which nutrients/elements are required for 96% of the plant (organic compounds)
C, H, O
Which nutrients/elements are required for amino acids specifically?
N, S
Which nutrient/element is required for regulation of stomates?
K
Which nutrient/element is required for integrity of the cell wall?
Ca
Which nutrients/elements are required for chlorophyll?
Mg, B, Zn
Which nutrients/elements are required for photosynthesis specifically?
Cl, Mn
Which nutrient/element is required for energy / ATP?
P
What can cause a deficiency in a plant?
Not enough of a mineral or restricted movement in plant.
What are the top three limiting nutrients?
Phosphate (P), Potassium (K), Nitrogen (N)
What does a phosphate-deficient plant look like?
Red/purple coloration on edges/margins of leaves.
What does a potassium-deficient plant look like?
Chlorosis - yellowing on edges/margins of leaves.
What does a nitrogen-deficient plant look like?
Chlorosis - yellowing in center of leaf between veins.
Chlorosis can lead to what?
Necrosis (leaf death)
Where do you see deficiencies when elements are mobile?
In old leaves (elements are drawn to sinks at new leaves)
Where do you see deficiencies when elements are immobile?
In new leaves
What does a Ca (immobile) deficiency in tomatoes produce? Why?
Tomato blossom end rot - cell walls are mushy
What is the ideal pH for soils?
Average around 7, but slightly acidic may be advantageous in some cases (higher H+ conc. can displace positively charged minerals from soil particles making them more available for absorption)
Why are heavy quantities of some nutrients detrimental?
Toxicity or overdoses can result and harm the plant.
What does nitrogen fixation do?
N2 --> (NH3 -->) NH4+
What does nitrification do?
NH4+ --> NO2- + NO3-
What does denitrification do?
NO3- --> N2
What is the preferred compound of nitrogen for the plant?
NO3- (can accept NH4+, but not preferred)
What do plant enzymes reduce NO3- to in side the plant? Why?
NH4+ for synthesis of amino acids / organic compounds
What is the N fixing bacteria that has a symbiosis with legume roots?
Rhizobia
What does rhizobia do for plants? Why does it rely on the plant?
Rhizobia fixes nitrogen, but needs nitrogenase from leghemoglobin (which is found in eukaryotes)
What are Diazotrophs? Example?
Free-living organisms that fix nitrogen into NH3 (cyanobacteria)
What form do rhizobium bacteria assume inside the vesicles in the root cell?
Bacteroids
What are the swelling in the legume roots? What causes these?
Nodules - caused by plant cells that have been "infected" by rhizobium
Each legume has a specific strand of Rhizobia, how do they recognize their specific strand?
Flavonoids - chemical signals
Which enzyme breaks the N2 triple bond to make NH4+? How much energy does this take? What is one specific requirement?
Nitrogenase
A significant amount (16 ATP)
Can't be exposed to O2
What is the protein that regulates O2 concentration so it is not harmful to nitrogenase?
Leghemoglobin (produced by eukaryotes, symbiosis)
Do Rhizobia + Legumes generate more or less usable nitrogen for plants than industrial fertilizers?
More
What is "Green Manure"?
Growth of crops just to produce nitrogen for the soil, plow it up into soil and then plant your harvesting crop.
Restores concentration of fixed N2 in the soil.
Rice has a symbiosis with what? How does this symbiosis work?
Azolla (fern with mutualistic cyanobacteria Anabaena, which fixes N2)
Rice kills the Azolla releasing nitrogen-rich organic material for increased fertility)
What are the two kinds of mycorrhizae? How are they different?
Arbuscular mycorrhizae (endomycorrhizae - penetrates cortex/cells making arbuscules - branched structures)
Ectomycorrhizae (encloses surface around roots)
What evolutionary characteristic was helpful in the colonization of land by plants?
Mycorrhizae symbiosis
What is an epiphyte? What kind of relationship does it form?
A plant that lives on the surface of another plant; gets water and nutrients from rain via leaves (not roots)
Forms a commensal relationship
What are four kinds of parasitic plants we talked about?
Mistletoe
Witchweed
Indian Pipe (ghost flower)
Dodder
What are carniverous plants?
Venus Flytrap and Sundews - obtain N from insect prey
What are the four main parts of an angiosperm?
Sepal, Petals, Stamen (m), Carpel (f)
Which part of the angiosperm is green and leaflike; it encloses and protects the floral bud?
Sepal
Which part of the angiosperm is brightly colored to attract pollinators?
Petals
Which part of the angiosperm contains the male reproductive portion? What two parts make up this structure?
Stamen (anther and filament)
Which part of the angiosperm contains the female reproductive portion? What three parts make up this structure?
Carpel (stigma, style, and ovary)
In the ABC theory, what do the genes (A, B, and C) make?
A = sepals
A+B = petals
B+C = stamens
C = carpels
Which part of the stamen contains the pollen sacs? Connects that part to the base of the flower?
Anther
Filament
Which part of the carpel is the landing pad for the pollen grain? Transfers the pollen tube? To the ___ containing the ovule?
Stigma
Style
Ovary
In the ABC theory, what is the rule of thumb?
A replaces C, C replaces A.
(ex: Lacking A, extra C (carpels),
Lacking B; just A and C;
Lacking C, extra A (sepals and petals)
What is characteristic of a complete flower?
It contains all four parts of an angiosperm.
What characterizes an angiosperm described as "imperfect"?
Only has one sex (stamens or carpels).
What are the steps for an angiosperm life cycle? (7 steps)
1) microsporangium contains microsporophytes that divide by meiosis to produce microspores (male)
2) microspore becomes pollen grain (male gametophyte) (n)
3) megasporangium produces megaspores (n) to form female gametophyte
4) pollination at stigma, pollen tube forms, two sperm cells enter ovule
5) double fertilization (sperm + egg = zygote (2n); sperm + polar nuclei = endosperm (3n)
6) zygote forms seed / fruit
7) seed germinates develops into mature sporophyte (2n)
The male gametophyte is made up of two cells...?
Pollen tube cell which contains generative cell
What does the generative cell do in the pollen grain?
Produces 2 sperm for double fertilization.
In the female gametophyte, there are how many nuclei? What are the names of all of them?
8 haploid nuclei:
- 3 antipodal cells
- 2 polar nuclei
- 2 synergids
- 1 egg
What is the function of the synergids?
They guide / attract the pollen tube to the egg.
What is pollination?
The transfer of pollen from anther to stigma for double fertilization.
In what ways can pollination occur?
- Abiotically by wind
- Pollination by bees
- By moths / butterflies
- By flies
- By birds
- By bats
What is meant by the "self-breeding depression"?
When flowers/plants are complete and self-fertilize leading to low genetic variation.
What is meant by an "incomplete" flower?
Missing any one part of the four angiosperm parts.
What is a "monoecious" plant?
Self-fertilized.
What is a "monoecious" plant?
Self-fertilized.
What is a "diecious" plant?
Only has male or female parts, not both; prevents self-fertilization.
What is a "diecious" plant?
Only has male or female parts, not both; prevents self-fertilization.
What is a Thrum flower? What is its function?
Short styles (carpels) and long stamens; makes it unlikely that a pollinator will get pollen to the stigma of the carpel.
What is a Thrum flower? What is its function?
Short styles (carpels) and long stamens; makes it unlikely that a pollinator will get pollen to the stigma of the carpel.
What is a Pin flower? What is its function?
Short stamens and long styles (carpels); makes it unlikely that a pollinator will obtain the pollen from the stamen.
What is a Pin flower? What is its function?
Short stamens and long styles (carpels); makes it unlikely that a pollinator will obtain the pollen from the stamen.
What are S genes?
Genes that recognize the self (or close relatives) and prevents self-fertilization.
What are S genes?
Genes that recognize the self (or close relatives) and prevents self-fertilization.
In double fertilization, where do the two sperm go to? To produce what?
Sperm + Egg = Zygote (2n)
Sperm + 2 Polar Nuclei = Endosperm (3n)
In double fertilization, where do the two sperm go to? To produce what?
Sperm + Egg = Zygote (2n)
Sperm + 2 Polar Nuclei = Endosperm (3n)
What does the ovule become?
The ovary?
Ovule => Seed
Ovary => Fruit
What does the ovule become?
The ovary?
Ovule => Seed
Ovary => Fruit
What are the steps leading from the zygote to the seed?
Cell division - unequal sized cells allows polarity determination;
Leads to embryo containing root apex and shoot apex;
Cotyledons stick out off of shoot apex end;
Integuments form hard seed coat.
What are the steps leading from the zygote to the seed?
Cell division - unequal sized cells allows polarity determination;
Leads to embryo containing root apex and shoot apex;
Cotyledons stick out off of shoot apex end;
Integuments form hard seed coat.
What are the four kinds of flowers/fruits? What distinguishes them from one another? (example)
1) Simple fruit - single or several fused carpels (pea)
2) Aggregate fruit - >1 separate carpels each forming small fruit (raspberry)
3) Multiple fruit - multiple flowers tightly clustered that fuse together (pineapple)
4) Accessory fruit - "fruit" forms from another part of plant, like enlarged receptacle, low in stem (apples)
What are the four kinds of flowers/fruits? What distinguishes them from one another? (example)
1) Simple fruit - single or several fused carpels (pea)
2) Aggregate fruit - >1 separate carpels each forming small fruit (raspberry)
3) Multiple fruit - multiple flowers tightly clustered that fuse together (pineapple)
4) Accessory fruit - "fruit" forms from another part of plant, like enlarged receptacle, low in stem (apples)
What are the three generations seeds contain?
- Mother sporophyte
- Gametophyte
- M/F new sporophyte
What are the three generations seeds contain?
- Mother sporophyte
- Gametophyte
- M/F new sporophyte
In what ways can seeds be dispersed?
Water, wind, animals.
In eudicots with thick cotyledons, what structures are present?
Seed coat, radicle, hypocotyl, two cotyledons.
In eudicots with thin cotyledons, what structures are present?
Seed coat, endosperm, cotyledons, hypocotyl, radicle.
What structures distinguish monocot seeds from eudicot seeds?
Coleoptile, coleorhiza, and pericarp.
What is the structure that covers the young shoot in monocot seeds? Covers the young root?
Coleoptile (shoot)
Coleorhiza (root)
What is the pericarp? What happens to it in monocot seeds?
Pericarp is the fruit / mature ovary tissue; it is fused to the seed coat in monocots.
What is the structure that stores nutrients that can be used by seedling after germination?
Endosperm
What are the structure(s) that absorb nutrients from endosperm and transfer to the rest of the embryo when seed germinates?
Cotyledons
What is the most commonly utilized herbicide (most kg applied)?
Herbicides
If a species is adapted to low resource availability, is it likely to have a smaller or larger seed?
Larger seed (more nutrients stored...)
When a seed stops growing and the metabolism nearly ceases, it is said to be?
Dormant
In what five ways can seed dormancy usually be broken / get seed to germinate?
1) Pre-chilling (common in middle latitudes due to winter --> spring)
2) Smoke can stimulate in fire-adapted species
3) Seed coat weakened by chemical attack (animal digestive tract)
4) Light stimulation (buried shallow enough)
5) Substantial rainfall
What is the first organ to emerge from a germinating seed? What does it form?
Radicle - forms root system
What occurs after the radicle/root system emerges?
Hook forms in hypocotyl; growth pushes the hook above ground.
When the (eudicot) hypocotyl hook pushes up out of the ground during germination, the cotyledons can be above ground or under ground, what are these called?
Cotyledons above soil = epigeal germination
Cotyledons below soil = hypogeal germination
Monocots' radicle emerges first during germination, followed by what?
Coleoptile pushes upward through soil and into the air (shoot system)
What is asexual reproduction?
Vegetative reproduction in plants; offspring from single parent; no genetic recombination; forms clones.
When fruits and seeds are produced int he absence of fertilization / pollination, it is called?
Apomixis
What is the main historical example of artificial selection in "plant breeding"?
Maize / Corn
What does it mean to be "transgenic"?
Genetically modified to express gene(s) from another species.
What is a plant cell that has had its cell wall removed (by cellulase and pectinase)? Why is this done?
Protoplast; for plant breeding.
How do scientists add plasmids to protoplasts (cells with a removed cell wall)?
Shoot protoplasts with gold beads with DNA on them; DNA from bacteria/gene is incorporated into DNA of plant.
What are the three steps in signal transduction?
1) Reception (detects signal from environment
2) Transduction (communication)
3) Response by plant
Where are the receptors for plants often located?
Trans-membrane molecules
What mechanism explains xylem movement? Phloem movement?
Transpiration = xylem
Pressure-Flow hypothesis = phloem
Hormones usually have what concentration in plants?
Low
What is the response of plants called when they bend towards light?
Phototropism
What is the primary function of Auxin (IAA)?
Cell elongation; enhances apical dominance; produced in apex, transported downward for root growth.
What is the primary function of Cytokinin?
Cell division; involved in apical dominance; produced in roots, transported up to shoots.
Which hormone is involved in stem elongation and growth and stimulates germination of seeds?
Gibberelins
Which hormone is a repressor of plant growth and is in the roots (when there is a drought it moves to leaves and tells guard cells to pump out K+ to close stomates).
ABA (Abscisic Acid)
Which of the plant hormones is a gas? It is involved in apoptosis and senescence (programmed cell death).
Ethylene
What is the onset of programmed cell death called?
Apoptosis
What is programmed cell death also known as?
Senescence
Which hormone is a signal in the fall for plant leaves to fall; fruits to ripen?
Ethylene
What is the triple response?
A response to an obstacle: plant stem bends, thickens and continues to grow horizontally.
What are phytochromes sensitive to?
Red light
How do you go from Pr --> Pfr? Reverse?
Hit with red light.
Hit with far-red light.
Pr --> Pfr --> ???
Seed germination, control of flowering, etc.
Which kind of plants will flower when there is a long night interrupted by a red light?
Long day, short night plants.
Which kind of plants will flower when there is a long night interrupted by a red light followed by a far-red light?
Short day, long night plants.
Which kind of plants assimilate CO2 better at low concentrations? At high concentrations?
Low = C4
High = C3