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137 Cards in this Set

  • Front
  • Back
Scientific fact
has occurred and is occurring, never accepted 100%
theory
processes by which evolution has occurred and continues to occur
Levels of evolution
1) Physical- origin of universe and Earth
2) Chemical- origin of life
3) Biological- origin of diversity
4) Cultural- origin of human social structure
Physical (levels of evolution)
origin of universe and Earth
Chemical (levels of evolution)
origin of life
Biological (levels of evolution)
origin of diversity
Cultural (levels of evolution)
origin of human social structure
Jean Baptiste de Lamarck
French, early 1800s- inheritance of acquired characteristics- disproven
Darwin
studied finches on Galapegoes Islands
On the Origin of Species
1859
 First printing sold out in 1 day
 Introduced concept of natural selection as mechanism of evolutionary change
 In 1900s, Mendel’s work rediscovered
 Modern Synthesis- 1940s
 Microevolution
 Macroevolution
Microevolution
adaption in populations
Macroevolution
evolution of new species
Darwin's theory
descent with modification
part 1: reproduction
part 2: variation
part 3: struggle for existence
part 4: natural selection
Continuous variation
o Continuous- numerous, intergraded phenotypes in population
 Bell curve
 Human height
Discontinuous variation
discontinuous

* many, discreet phenotypes in population
* no pattern seen in graphs
* human ABO blood types
Struggle for existence
* more offspring produced than can be sustained
* members of a population show variation
* must compete for limited resources and mates
* what decides which will compete successfully?
Natural Selection
* organisms whose genotypes result in a phenotype that better adapts them to their environment
o beneficial mutations or advantageous gene combinations
* reproduce and add disproportionately to gene pool of next generation
o compete for /utilize resources and space
o avoid predators and disease
o attract mates
o develop advantageous associations: mutualisms
artificial selection
used by darwin as a model for natural selection with pigeon breeding
Herbert Spencer
survival of the fittest
_______ acts as a filter of natural selection
environment

natural selection:
* responds to:
o immediate environment
o predictable environment
* cannot plan for unusual or long term events
* change in environment changes favored variations
o what is adaptive in one environment may not be in another
"Selection for"
greater reproductive potential
"Selection against"
less reproductive potential
Stabilizing Selection
* average or mean phenotype is selected for and the extremes are selected against
* classic example:
o extreme low birth weight babies and extreme high birth weight babies have lower survival rates (i.e., are selected against); average weight babies have higher survival rates (i.e., are selected for)
* effect: average phenotype stays the same over time
Directional Selection
* one extreme are selected for or against
* classic example:
o peppered moths (Biston betularia)
o two forms: gray and black
+ gray form better camouflaged on trees from hungry birds
o before 1840’s in England - gray more common
o after 1840’s, Industrial Revolution
o trees turned black with soot, black form now camouflaged and became the more common form
* effect: average phenotype shifts directionally
Diversifying or Disruptive Selection
* average phenotype is selected against and the extremes are selected for
* example:
o African butterflies
o members of a widely dispersed, tasty species will locally mimic various other unpalatable species
o within the population there are several phenotypes
* effect: the population shows a variety of distinct phenotypes, with a least a bimodal distribution
Hardy-Weinberg Theorem
* measure of genetic change in a population

# assumes:

* no selection
* no mutation
* no migration
* large population
* random mating

# predicts:

* no change in gene frequency in population
Population Change
* If change is seen, one or more H-W assumptions must be violated
* Evolution: change in gene frequency in a population over time
* population phenomenon
o individuals cannot evolve
Speciation
evolution of a new species

origin of isolating mechanisms
species
a group of actually or potentially interbreeding populations that cannot interbreed with other such groups
prezygotic
prevent mating - examples:

* geographical - separated by geographical barriers
* mechanical - genitalia do not fit
* behavioral - different mating behaviors
postzygotic
offspring cannot reproduce

* hybrids sterile (mule)
* hybrids inviable - embryos abort (goats and sheep)
* hybrid breakdown - next generation not viable
Allopatric
* geographical barrier separates populations
* no interbreeding
o each with unique mutations
o each facing unique selection
* over time --> diverge and cannot interbreed
Sympatric
* new species arises without geographical isolation
* new species in a single generation
* polyploidy - doubling of chromosome number
o irregular mitosis or meiosis
o can only interbreed with itself
o common in plants
Adaptive Radiation
* proliferation of species from a common ancestor by adaptation to various niches
* small founder population invades a new territory
o few competitors
o ==>variation, new niches
o ==>inbreeding and isolation
o ==>speciation
* Darwin’s finches on Galapagos Islands
Phylogeny
* evolutionary lineage
* tracing a phylogeny
o biogeography -geographic distribution
o paleontology - fossil record
o comparative anatomy
+ homologous organs - similar due to common descent
+ analogous - similar function, different origin
o molecular homology
o chromosomal homology
Earth is...
4.6 billion years old
Universe is...
14-15 years old
Oldest fossil
3.6 billion years old
bacterialike
eukaryote
earthly atmosphere in beginning
methane, nitrogen, hydrogen, ammonia, water vapor

no oxygen
Miller
sparks in experimental atmosphere

1 week --> amino acids, sugars; nucleic acid bases, lipids found by others
Origin of Life: Membranes
* lipids and proteins will spontaneously form semipermable membranes
* form into enclosed droplets called coacervates
* coacervates will “grow” and “multiply” although this only mimics living systems
* may be a prototype for very early pre-cells
stumbling blocks
* polymerization of amino acids to protein
o may occur on hot wetting/drying surfaces
o may occur on clay crystals(life from dust of Earth?)
o controversial
* polymerization of nucleic acids and origin of genetic code
o possibly the first genes were RNA, not DNA
Origin of Life
RNA trapped in protocell membrane may have been first (VERY simple) cell
Origin of Life - Mars
* Recent discoveries in Martian meteorite
* 3 lines of evidence
o organic material in shape of bacteria
o iron residues in form associated with life
o PAH residues associated with life, clustered on the rock
sexual reproduction
mitosis and meiosis
asexual
mitosis only
animal life cycle
gametes, fertilization, zygote, mitosis, development/growth, adult, gonad, meiosis
animals: some organisms use both _____ and _____ reporduction
sexual and asexual
sperm is produced in the _____ by ______
testis (male) and ovary (female) by meiosis
oviducts
carry eggs from ovary
uterus
site of embryonic devlopment
hermaphrodite
one individual with both sex organs

earthworm, sea slugs
external transfer of sperm to egg
usually in equatic animals
sperm and egg released into environment
internal transfer of sperm to egg
sperm placed in female tract

variety of mechanisms
- sperm packet for spiders
- penis
Development
egg and sperm, fertilization, zygote, embryo
fertilization
- fusion of one sperm and egg
- activates development process
- egg becomes instantaneously unreceptive to other sperm
zygote
* single cell from fusion of egg and sperm (by definition)
* sperm contributes only nucleus to cell, all other sperm cell structures degraded by zygote
* cytoplasmic organelles derived from egg only, i.e. from mother
o trace mitochondrial DNA to original modern human population
o possible African origin 200,000 years ago
development (animal reproduction)
* zygote begins to divide by mitosis
o cleavage
* ball of cells same diameter as egg
o subdivides cytoplasm
* embryo
o from 2 cells onward in early development
o nutrients provided by female parent
o develops by mitosis
* solid ball of 16 cells = morula

gastrulation
* invagination form at one point on hollow sphere
* like a finger poking into a balloon

* invagination reaches opposite side and fuses
* hollow tube through sphere = gastrula
blastula
hollow ball of cells
gastrulation
* invagination form at one point on hollow sphere
* like a finger poking into a balloon
archenteron
archenteron
endoderm
inner tube
ectoderm
outer sphere
openings (development)
mouth and anus
ectoderm (development)
skin and nervous tissue
mesoderm (development)
muscle, bone, other organs
endoderm (development)
lining of gut, lungs, other digestive organs
gastrula (development)
growing embryo with developing organs
Overall of development
zygote, cleavage, morula, blastula, gastrulation, gastrula, 3 germ layers, developing embryo
male gonads =
testes, in scrotum
paired, external to body cavity
sperm production is temperature sensitive
varicocoele
varicose veins in scotum
raises temp and reduces sperm count
epididymis
stores sperm
adjacent to testis
vas deferens
sperm duct
over bladder and to urethra
paired
vasectomy = cut and tied
urethra (male)
common duct for urine and semen
penis
copulatory organ;
shaft, prepuce (foreskin), glans
rich supply of nerve endings
glands (glands) (sets)
add fluid to semen
3 sets:
- seminal vesticles
- prostate gland (majority of fluid), prone to cancer
- bulbourethral glands
Spermatogenesis
* occurs in seminiferous tubules of testes
* 1 mother cell --> 4 sperm
o meiosis and sperm maturation 65-75 days
o continuous from puberty onward
* fertility
o fertile: 100 million sperm/cc
o infertile: <40 million sperm/cc
o sterile: no sperm present
infertility (male)
low/no sperm count
causes of infertility/sterility

* varicocoele-varicose veins within the scrotum
o raised temperature around testes
o repaired by tying off some veins to reduce blood flow
* vasectomy
o cut and tie vas deferens
* radiation, chemicals
o cancer treatment
Semen
mix of sperm (5%) and glandular fluids (95%)

alkaline to counteract acidity of vagina
Steps of Ejaculation
* Stage 1
o rhythmic contractions of epididymis, vas deferens, prostate
* Stage 2
o rhythmic contractions of urethra and surrounding muscles
* orgasm very stereotyped in males
o tends to occurs in a similar fashion in all males
Hormonal Control-Male
Before birth
* human chorionic gonadotropin (HCG) from placenta stimulates testes to produce testosterone

---> primary sex characteristics:

* genitalia and sex organs

at birth, testes become dormant for 10-15 years
Testosterone- secondary sex characteristic
* genitals enlarge
* voice deepens
* beard and body hair
* baldness
* muscles enlarge - anabolic effect
Anabolic Steroids
* illegal- used to increase muscle mass
* analogs of testosterone
* must be in greater than ambient amounts for “desired ”muscle enlargement
o inhibition of testosterone and sperm production
o testicular atrophy (testes shrink in size)
o body organ damage (esp. liver and kidney)
o injected
+ shared used of syringe can transmit HIV
+ uncertain origin of drug-suppliers not pharmacists!
ovaries=
gonads
* paired, in body cavity, 1 in. long
* produces hormones and eggs
* connective tissue and primary oocytes
* eggs develop to prophase I before birth and “freeze” at that stage until ovulation
fallopian tubes =
oviducts
* transport egg to uterus
* site of fertilization
* open into body cavity
* fimbrae surround ovary and sweep egg into oviduct
* tubal ligation
o fallopian tubes cut and tied
o renders woman sterile
uterus
# muscular sac (smooth muscle)
# above bladder
# inner lining = endometrium

* enriches with blood vessels for pregnancy
cervix
* opening (os) from uterus to vagina
* diam. of a pencil
* entry of sperm; birth of baby
* filled with mucous that thins at ovulation to permit sperm to enter - protection against bacteria
vagina
* receive penis (semen) in copulation
* birth canal
* contains glands for lubrication at copulation
external genetalia
vulva

* mons pubis
* labia majora - external
o contain erectile tissue
o homologous to scrotum
* labia minora - internal to labia majora
o surround vaginal opening
vulva
* clitoris
o homologous to glans
o rich supply of nerves
* urethral opening
* hymen
o covering over vaginal opening
o may be ruptured in first intercourse or sports, etc.
female sexual response
- labia majora becomes erectile
- vagine secretes lubricant
- orgasm
stages in both sexes
1) excitement
2) plateau
3) orgasm
4) resolution
oogenesis
1 mother cell --> 1 functional egg
ovulation:
day 14
basal body temperature increases about .5 C
body temperature upon waking
basis for natural family planning

ovulation kit measures levels of LH
LH Surge
Day 13
pregnancy
implantation in endometrium on day 19-21
4-7 days from fertilization to implantation
embryo forms early placenta
pregnancy steps
corpus luteum, prosterone, maintains, embryo-placenta, HCG
early embryo (human development)
blastocyst
5 days at implantation
inner mass, embryo proper
trophoblast, chorion and placenta
extraembryonic membranes
part of embryo

o chorion = outer sac, from trophoblast
o amnion = inner sac, from inner mass
o contains fluid = amniotic fluid
+ some fetal cells in fluid
+ removal for amniocentesis
embryo structures:
allantois
yolk sac
umbilical cord
* allantois
o early kidney
o from inner mass
* yolk sac
o stores germline cells
o from inner mass
* umbilical cord
o formed in part from all extraembryonic structures
o connects at abdomen --> navel scar
placenta (associated with embryo)
* interface between mother and embryo
* exchange of nutrients, gases, and wastes
* fetal and maternal blood separated by membranes, do not mix
* drugs, antibodies, viruses but not bacteria cross placenta
o cuation should be used with all drugs beucase they can cross the placenta and possibly affect the developing embryo/fetus
first trimester
implantation
organogenesis (organs form)
embryo very susceptible to drugs and disease
second trimester
embryo becomes fetus
ossification of bones
movement
organ maturation
third trimester
period of growth
progesterone and estrogone trigger production of relaxin and prolactin in mother
relaxin
loosens ligaments and allows pelvis to open for birth
prolactin
stimulates milk production
birth
fetus moves w/ head at the cervix
three stages:
1) contractions- 'water breaks'
2) expulsion of baby
3)expulsion of placenta
sexual reproduction (plants)
# overall very similar to animals but with a few important differences
# details used in classification of plants
# commercially important

* alternation of multicellular haploid and diploid phases in the life cycle
* diploid phase called sporophyte
* haploid phase called gametophyte
* Example: the fern
asexual reproduction (plants)
# vegetative
# commercially important
diploid phase
sporophyte
haploid phase
gametophyte
sporophyte
# 3 and 4 in the fern life cycle diagram
# diploid phase, often the most conspicuous phase

*fern plant, geranium plant, pine tree, oak tree

# part of the plant becomes specialized structures called sporangia in which meiosis occurs

* in ferms, looks like fuzzy brown hairs on underside of leaf

# meiosis produces spores in plants
# spores are haploid, single-celled and often have a thickened cell wall
# “sporophytes produce spores in sporangia”
Gametophyte
# 1 in the fern life cycle diagram
# haploid phase, usually very inconspicuous or hidden

* fern gametophytes are about the size of the head of a pin

# spores are the first stage in the gametophyte phase
# produce gametes, eggs and sperm, in specialized structures by mitosis
# “gametophytes produce gametes”
angiosperm life cycle
involves flower, fruit and seed
Flower
# modified stem tip with modified leaves
# very diverse in appearance but all flowers will have the same basic structures
# flower is an organ for sexual reproduction
5 basic parts of a flower
recepticle
sepals
petals
stamens
pistil
pistil
# carpel = basic unit

# structure in which seeds are produced
# parts:

* stigma - receives pollen
* style - connection between stigma and ovary
* ovary - closed container of ovules/seeds

o ovule = unfertilized seed
* develops into the fruit
stamens
produce pollen (in anther)
petals
- colored, often showy
- typically for attracting pollinators
sepals
- green
- cover and protect the flower
recepticle
base on which other parts are attached
pollen is ____ and stigma is _____
male; female
hay fever
allergic reaction to the proteins in the pollen
pollination
transfer of pollen from anther to stigma
self compatibility
can self pollinate
self-incompatibility
cannot self pollinate
mechanisms for pollination
wind and animals
ovule
develops in ovary

- when fertilized, becomes the seed
-structure: outer coat, inner cell mass, within which a cell becomes the megaspore mother cell
sporangium
inner mass of cells in ovule, female
female gametophyte (ovule in angiosperm life cycle)
egg and central cell with two haploid nuclei
fertilization
first: pollination, then pollen germinates into a pollen tube

pollen tube grows into style and to ovary and to the ovule

tube carries two sperm
double fertilization
unique to flowering plants

sperm, egg, zygote, mitosis, embryo

other sperm, central cell fuses with the two haploid nuclei, endosperm nucleus, mitosis, endosperm

(endosperm is a nutritive tissue)
embryonic development (angiosperm life cycle)
zygote divides into an embryo
- embryonic root
- embryonic shoot
- cotyledons
Seed part
seed coat
embryo
stored food (endosperm)
fruit and functions
mature and ripened ovary,
development signaled by hormones from seeds

functions:
- seed protection
- seed dispersal
types of fruit
fleshy- soft outer layer
dry- hard outer layer
Asexual reproduction in plants
mitosis only- no meiosis
- very common
- commercially important
- cloning