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94 Cards in this Set

  • Front
  • Back
symbiotic associations
organisms of different species living together
Facultative symbiosis


Obligate symbiosis
-Symbionts can exist independently of one another, both can still benefit (clown fish & sea anemone)

-one or both organisms must be together to survive (termite & organism in body)
coevolutionary relationships
symbionts interact and evolve evolutionarily
Mutualism
Both symbionts benefit
>coral (oxygen, glucose)+zooxanthellae (protection, CO2, home)
>ants (honeydew)+aphids(protection)
>hermit crabs (protection from predators)+ sea anemones (motility)
Parasitism
Parasites benefit, host is harmed
-necrotrophic= host is eventually killed by parasite
-biotrophic= host lives with parasite (most parasites have evolved to be biotrophic)
-ectoparasites= live outside host
-endoparasites= live in hosts
>flea, bed bugs
>cuckoo + warbler
>oyster pea crab+oyster
-parasitoid= parasite lives in/on host while developing, kills host once development finishes, becomes free-living
>wasp+catalpa worm (caterpillar of sphinx moth)
Commensalism
One animal benefits, other neither harmed nor benefits
>Sloth+ sloth moth (eats algae on sloth fur)
-phoresy= one animal uses another only for transportation
>bird louse+louse fly
-inquilinism= one animal uses another for housing
>barnacles+whales
-matabiosis= one animal takes advantage of suitable environment provided by another
>hermit crab+snail shell
Amensalism
one symbiont harmed, other unaffected
-competition
>mussles living on a substrate with other filter feeders
Ethology
study of animal behavior, especially under natural circumstances
Proximate Causation
how an animal triggers a behavior, immediate physiological causation of a behavior
Ultimate Causation
why an animal does a behavior
taxis
response to a stimulus by moving toward or away from it
-phototaxis=response due to light
-thermotaxis=response to temperature
-thigmotaxis=response to physical contact
reflex
involuntary response to a stimulus
Greylag goose study
female sees egg out of nest, rises from nest, uses head/neck/bill to retrieve egg
-completes task in orderly fashion even if something changes, must go all the way through sequence
-fixed action pattern= stereotypical behavior that is a response to a situation
-releaser/sign stimulus= stimulus that initiates behavior (can produce wrong behavior)
innate behavior
inherited behavior, do not have to learn behavior to survive and reproduce
Habituation
innate response to a stimulus decreases
-low intensity stimuli, non-painful
>police horse in a big city
Sensitization
innate response to a stimulus becomes larger than usual, increases
-high-intensity stimulus, painful and unpleasant
>aquatic snail withdraw gills with electric shock
Imprinting
specific to young animals, cause by exposure to certain stimuli during correct part of development
>ducklings& goslings follow first large object they see
Societal Learning/ Learning in a Society
learning from a social group
>macaque washing and eating food
Animal communication
used to influence behavior of another animal
-signals can be sound, odor, movement, touch, visual...
Social behavior
any interaction that results from a response from one animal to another of the same species
-brief/temporary or long range
>competition (food, mates, territory, any necessity)
>territoriality (territory= fixed area whose occupant keeps out intruders of same species)
>mating systems (monogamy= one male and one female pair up for life or breeding season, polygamy= either male or female has multiple mates)
polygyny= 1 male+multiple females
polyandry= 1 female+multiple males
integument
protective outer-body covering, semi-permeable so it can regulate material internally/externally (water balance, temp balance)
-Inverts: epidermis (epithelium), non-cellular cuticle secreted by epidermis
-Verts: epidermis (stratified squamous epithelium), dermis (dense fibrous connective tissue)
Keratinzation
as epithelial cells age, they move up= take in keratin for protection, eventually shed or become removed
skeletal system
many skeletal systems are not composed of bone
function: protection, structure, movement, site of muscle attachment
hydrostatic skeleton
fluid-filled cavity used for structure and movement
-fluid puts pressure on integument to give structure, allows for muscle function
>earthworms, cnidaria, nematoda, sea stars
muscular hydrostat
lacks bone and fluid support, muscles themselves provide support
-muscles are non-compressible
>mammal tongue, elephant trunk, octopus tentacle
rigid skeleton
rigid parts for muscles to attach to
-muscles work in opposing sets= extensors& flexors, muscles only pull so they need an attachment place
-rigid exoskeleton= outside body, for protection & locomotion & structure, secreted by integument (will grow with animal) or integument is rigid exoskeleton (must be molted=ecdysis)
>insects, inverts, molluscs, clams, crabs
-rigid endoskeleton= inside body, locomotion & structure, can grow with animal, calcareous plates or bones/cartilage
>sea stars, sea urchins, verts
Bone
-spongy/cancellous bone= light, porous, mesh work of materials
trabecula= rod shape, form lattice work of bone where most stress is for max strength
pores filled with marrow= produce erythrocytes and leukocytes (found in ends of long bones or flat bones)
-compact bone= more dense, more calcium, more matricx, less porous, appears solid
osteon= outer layer of all bones, most of shaft of all bones
Muscle
-fascicle=bundle of muscle fibers surrounded by connective tissue
-muscle fibers= cells that make up fascicle
-myofibrils= make up muscle fibers, run lengthwise
-sarcolemma= plasma membrane around muscle fibers, extend into cells to connect myo...
-fascicle=bundle of muscle fibers surrounded by connective tissue
-muscle fibers= cells that make up fascicle
-myofibrils= make up muscle fibers, run lengthwise
-sarcolemma= plasma membrane around muscle fibers, extend into cells to connect myofibrils
-sarcoplasmic reticulum= mesh-like organelle surrounding myofibril, calcium storage
-sarcomeres= sets of overlapping protein filaments in repeating patterns that make up myofibrils
>z-lines= bands that separate each sarcomere
-actin&myosin= main proteins of sarcomeres, responsible for muscle contraction
>troponin&tropomyosin are secondary proteins around actin filaments
Calcium in Muscle Function
calcium is released from sarcoplasmic reticulum, binds to troponin and changes its shape, confirmational (shape) change pushes tropomyosin off of resting place to reveal active sites on actin filament, club-shaped heads of myosin attach to active sites of actin and pull actin along= sarcomere shortens and muscle contracts
motor units
motor neurons + all muscle fibers/cells in innervates
-multiple motor units per muscle
-more motor units recruited=more muscle force
-fine muscle control

(motor neurons stimulate muscle fibers to contract, stimulate movement)
Skeletal muscle fibers
-slow oxydative
-fast oxydative
-fast glycolytic
1. Slow oxydative fibers (slow twitch muscles)
-muscles that need to be active continuously
-slow, sustained contraction, fatigue resistant
-muscles high in these fibers= "red muscle"= highly vascularized, lots of mitochondria, ,yoglobin
2. Fast oxydative fibers (fast twitch muscles)
-fast, powerful contractions
-fatigue-resistand
-high blood supply, lots of mitochondria, lots of myoglobin= "pink muscle"
3. Fast glycolytic fibers (fast twitch muscles)
-fast, powerful contractions
-fatigues easily
-fibers are large in diameter=most myofibrils=most force
-whitish in color=less blood supply, less mitochondria, less myoglobin (function araerobically= without oxygen)
Phylum Porifera
Phylum Porifera
"Porifera"= pore-bearing
~15000 species, mostly marine, some fresh water
cessile=stay in one place
suspension/ filter feeders
depend on water currents for food
blastula stage of development= animal classification
Porifera body plan
-Cellular level organization (specialized cells to perform different body functions)
-no germ layers
-no coelom (no mesoderm)
-asymmetrical
-no cephilization
-no segmentation
-neither protostome nor deuterostome
Porifera structure
Pores
-ostia/ostium= incurrent pores, allow water into sponge
-oscula/osculum= outcurrent pores, larger than ostia
spongocoel= cavity in some sponges
endoskeleton
-mesh work of extracellular material=scaffolding for living cells
-secreted by special cells
-spicules, help provide support, protection against predators, needle-like structure (Calcium carbonate, silica)
-collagen fibers= spongin
mesohyl=jellylike matrix in which spicules, spongin, and cells are imbedded
Porifera cells
-pinacocyte= flattened cells, line external surfaces and some internal surfaces, protective function, regulate pore size
-archaeochye (amoebocyte)= amoeboid cells. move around in mesohyl, digest and distrubute food (intracellular digestion, can b...
-pinacocyte= flattened cells, line external surfaces and some internal surfaces, protective function, regulate pore size
-archaeochye (amoebocyte)= amoeboid cells. move around in mesohyl, digest and distrubute food (intracellular digestion, can become any other type of cell if needed)
-choanocyte (collar cell)= line canals and chambers, flagella keep water flowing through sponge, food particles stuck to microvilli of collar
Porifera canal systems
-asconoid
-syconoid
-leuconoid
1. asconoid= simple, small, tube shape
-ostia>spongocoel>osculum
2. syconoid= tube or vase shape, thicker body
-ostia>prosopyles>radial canal>apopyle>spongocoel> osculum
3. leuconoid= most common, most complex, larges, irregular shapes, no spo...
1. asconoid= simple, small, tube shape
-ostia>spongocoel>osculum
2. syconoid= tube or vase shape, thicker body
-ostia>prosopyles>radial canal>apopyle>spongocoel> osculum
3. leuconoid= most common, most complex, larges, irregular shapes, no spongocoel
-ostia>incurrent canal>flagellated chamber> excurrent canal> osculum
Porifera reproduction
asexual
-fragmentation
-budding
-producing gemmules= internal buds (collegction of archaeocytes embeded in spongin of dying sponge)

sexual
-monoecious= male and female parts in one structure
Phylum Cnidaria
Phylum Cnidaria
"knide"= stinging cell
~9000 species
Body Plan
-Tissue level organization (no organs, beginning of nerve and muscle activity)
-doploblastic (lack mesoderm)
-no coelom
-radial symmetry (oral-aboral axis)
-no cephalization
-no segmentation
-neither protostome nor deuterostome
Cnidaria:

cnidocytes
specialized cell with stinging organelle, prey capture and defense, concentrated at tentacles
-nematocyst= stinging organelle
-cnidocil=trigger cilium, signals nematocyst to  be released
specialized cell with stinging organelle, prey capture and defense, concentrated at tentacles
-nematocyst= stinging organelle
-cnidocil=trigger cilium, signals nematocyst to be released
Cnidaria structure
-epidermis= outer layer, ectoderm, epitheliomuscular cells with dispersed cnidocytes and nerve cells
-gastrodermis= endoderm, nutritive-muscular cells (digestion, muscular contraction)
-gastrovascular cavity= incomplete gut/blind gut (only mouth...
-epidermis= outer layer, ectoderm, epitheliomuscular cells with dispersed cnidocytes and nerve cells
-gastrodermis= endoderm, nutritive-muscular cells (digestion, muscular contraction)
-gastrovascular cavity= incomplete gut/blind gut (only mouth, no anus)
-mesoglea= gelatinous, extracellular layer between epidermis and gastrodermis, secreted by epidermis cells, structural support
-nerve function= no central nervous system= nerve net=base of epidermis and gastrodermis on either side of mesoglea, network of cells
Cnidaria
-alternation of generations
dimorphic= 2 different adult forms
-polyp=asexual adult, sessile, specialized roles, often fixed on substrate, reproduce by budding
-medusa= motile adult, sexual reproduction, dioecious
>bell/unbrella shape, body parts in 4, tentacles, floating/weak swimming
>mouth at end of manubrium (stock)
>produces gametes

*some species have lost dimorphism and are either polyp or medusae
Phylum Cnidaria
Class Hydrozoa
Phylum Cnidaria
Class Hydrozoa
Colonia
-Genus Obelia= polyp form dominates, connected by coenosarc (epidermis, mesoglea, gastrodermis, shared gvc)
>perisarc=protective covering around coenosarc, made of chitin
>zooid= each individual polyp, hydranth= feeding polyp, gonangium= asexual reproductive polyp= free swimming medusae

Solitary
-Genus Hydra= only polyp formm sexual/asexual
>hypostome, basal disk/pedal disk
Phylum Cnidaria
Clas Scyphozoa
Phylum Cnidaria
Clas Scyphozoa
True jellyfish
-thick mesoglea, medusa form dominates, short-lived or no polyp stage
Phylum Cnidaria
Class Anthozoa
Phylum Cnidaria
Class Anthozoa
Sea anemone & coral
Largest class
Phylum Platyhelminthes
Phylum Platyhelminthes
"Flat worm"
~20000 species
Body plan
-organ level organization
-triploblastic
-acoelomate (mesenchyne="solid tissue layer)
-bilteral symmetry (dorso-ventrally flat)
-cephilization
-no segmentation
-protostome
Platyhelminthes structure
Incomplete gut= gvc= undigested food back through mouth
Nervous system
-ganglia= nerve cells in head with nerve cords running length of body
Excretory system
-protonephridia= tubules that excrete ammonia and maintain water balance
-flame cells= flagellated cells that keep fluids moving
Platyhelminthes reproduction
Sexual and/or asexual
Monoecious= hermaphroditic= male and female parts in one individual
Internal fertilization
Phylum Platyhelminthes
Class Turbellaria
Phylum Platyhelminthes
Class Turbellaria
Free-living flat worms
Marine, fresh-water, damp terrestrial environments
Mostly carnivorous
Muscular pharynx= extend out of body to seize prey
Eye spot= sense light
Auricles= sensing dissolved chemicals, smell
>planarians
Phylum Platyhelminthes
Class Trematoda
Parasitic flukes
-endoparasite= live in host body (mostly vertibrates)
Similar structure to turbellarians, except:
-no eye spot
- oral sucker
-hooks to hold on to host
Clonorchis sinensis

(Phylum Platyhelminthes, Class Trematoda)
Clonorchis sinensis

(Phylum Platyhelminthes, Class Trematoda)
Human liver fluke
-egg shed in human waste
-intermediate host: aquatic snail
Eggs hatch in snail, become miracidium (motile, ciliated), travel through snail tissue to become sporocyst (loose cilia), sporocysts produce multiple redia (oral sucker, germ cell ball), redia move to snail liver and mature into cercaria
-intermediate host: carp or minnow
Cercaria finds fish host and burrows into muscle tissue, loose tail and encyst, cercaria becomes dormant metacercaria, fish consumed by human raw or undercooked
-definitive host (where sexual reproduction occurs): human
Cyst dissolved in intestine= releases fluke that moves to liver to reside
Phylum Platyhelminthes
Class Cestoda
Tapeworms
Endoparasites= 2 hosts
Adults live in digestive tracts of vertibrates
No mouth or digestive system=absorb nutrients through body wall from host
-Cuticle= prevents from being digested
-Proglottids= reproductive units insegments attached in chain, continuously produced behind scolex, mature as they move down chain (end of chain= gravid= full of eggs)
-Scolex= unit of attachment with hooks and sucker (not a head)
Self-fertilize= one proglottid fertilizes another proglottid in same tapeworm, can cross-fertilize if another present
Taenia saginata

(Phylum Platyhelminthes, Class Cestoda)
Beef tapeworm
Human intestine, can live as longas host
Up to 10 meters
-eggsshed from gravid proglottid onto ground, consumed by cows, eggs hatch in cow, larvae move into blood vessels, embed and encyst in skeletal muscle (cstercus/cysterci), developes inside-out scolex in cyst, beef consumed by human raw or undercooked ("measly beef"), move into intestine and cyst wall dissolve, scolex turns right way and attaches to intestine wall, begins producing proglottids
Phylum Nematoda
Phylum Nematoda
Round worms= cylindrical worms with pointed ends
"Nema"= thread
Most less than 2.5mm long
About 2000 known species
Body plan
-organ level
-triploblastic
-pseudocoelomate (filled with fluid to distribute nutrientsand oxygen)
-bilateral symmetry
-cephilization
-no segmentation
-protostome
Nematoda structure
Complete gut= mouth and anus= digestion occurs in one way pth, specialization of gut
Cuticle= made of collagen, tough extracellular layer for protection
-ecdysis= shedding/molting of cuticle when outgrown
Only longitudinal muscles= can only move in thrashing motion
Nervous system
-ganglia
-dorsal and ventral nerve cord
-sensory organs clustered in head region
-nerves don't have axons= muscles have cytoplasmic extensions that send message to neurons= muscle arms
Each animal in each species has the exact same number of cells= eutely
Nematoda reproduction
Dioecious= males generally smaller than female, males have hooked end
-copulary spicules= on hooked end of male, aid in holding on for internal fertilization
Direct development
-ecdysis of cuticle between stages
-no true larval stage
Nematoda life cycle
Ascris lumbricoides
Intestineal round worms of humans (parasitic)
Adults live in small intestine & follow after food bolus (clump of food)
-eggs shed in feces, eggs have tough protective shell, juvenile develop in shell
-human consume soil or unclean plant material
-juvenile hatch in large intestine, burrow through intestinal wall into blood stream to lungs, get coughed up and swallowed again to reach small intestine
Nematoda life cycle
Soil-living= beneficial or parasitic, aid or destroy plants, can eat crop-consuming insects or other nematodes
Nematoda life cycle
Hook worm
Parasitic
Common in tropics
Adults up to 11mm long
Prevalence in US declined by use of shoes
-burrow into skin, make way to lungs, eventually make way to small intestine where they attach and suck blood

(Most nematodes are harmless to people)
Phylum Mollusca
Phylum Mollusca
soft body
marine, fresh water, terrestrial
Over 90000 species (about 70000 in fossil record)
Body plan:
-organ system organization
-triploblastic
-eucoelomate (small coelom around major organs)
-bilateral symmetry
-some cephalized, some not
-no segmentation
-protostomes
Mollusca form and function
-head-foot= ventral part of animal, feeding, locomotion, sensing environment
foot= very muscular, modified for each species
-visceral mass= contains most organs
-complete gut
-radula= feeding organ, rasping (scraping), rows of backward facing teeth made of chitin
(unique to mollusks)
teeth mounted on cartilage base= odontophore
teeth replaced by new ones when worn out
radula can be modified
-mantle= 2 folds of skin that are outgrowths of the body wall, protect visceral mass
-valve= shell (in some molluscs), secreted by mantle, more protection for visceral mass
-ctenidia= gills in mantle cavity, often ciliated
terrestrial molluscs have lungs rather than ctenidia
-mantle cavity= out spot for waste and reproductive products
Mollusca nervous system

Mollusca circulatory system
Nervous
-multiple pairs of ganglia connected by nerve cords
-sensory receptors (mechanoreceptors, chemoreceptors, statocysts, photoreceptors)
>photoreceptors= ocelli (sense light vs. dark) to eyes, or none at all

Circulatory:
-open circulatory system= blood not always confined to blood vessels, can travel through open sinuses in body
Mollusca reproduction
dioecious= separate male/female
direct and indirect development
Phylum Mollusca
Class Polyplacophora
Phylum Mollusca
Class Polyplacophora
"many plates"
>chiton
dorsoventrally flattened
mantle cavity runs on either side of foot
-ctenidia run on both sides
shell made of 8 curved plates
marine (rocky intertidal zone)
herbivorous= eat algae off of rocks
-veiled chiton eat invertibrates
Phylum Mollusca
Class Scaphopoda
Phylum Mollusca
Class Scaphopoda
tooth/tusk shells
conical shells open at both ends, mantle fused in cone shape to inside
marine (subtidal zone to depths)= burrowed into sand
not commonly seen
Phylum Mollusca
Class Gastropoda
Phylum Mollusca
Class Gastropoda
"stomach foot"
>snails, slugs, nudibranchs, abalone, limpits
most are herbivores, some carnivores, some detritovores
if they have shells, univalve (one shell)
-some coiled, shield like
size: microscopic to 1m
70000 species
cephalization, most have eyes (some on eye stocks)
locomotion: muscle contracts in foot
pulmonate=having a lung, lost gills
>snails & slugs in terrestrial and freshwater
-slugs= vestigial shell (no real purpose, not evolutionarily gone) or no shell, lung in mantle
highly vascularized lung= opens to pneumostome (air hole)
anus opens through pneumostome
partially detorted
Phylum Mollusca
Class Gastropoda torsion
torsion= 180 degree counterclockwise rotation of visceral mass, mantle, and mantle cavity
-occurs during larval stage
-bring mantle cavity to front of animal
-anus and mantle cavity open up over head (fouling= waste from anus can wash back over gills
-detorsion= partial return to untorsion state
-shell typically asymmetrically coiled
planospiral shell= symmetrical, lined up w/ body mass
conispiral shell= whorls go off to side, shifted to side (lost right ctenidium, kidney, atrium, reduce fouling)
Phylum Mollusca
Class Bivalvia
Phylum Mollusca
Class Bivalvia
two shells
>clams, oysters, mussels, scallops, ship worms
mostly marine, some fresh water
no cephalization, no eyes, some have ocelli
laterally compressed
mantle attached to valves (shells) and drapes over visceral mass
valves hinge together w/ elastic ligament, teeth hold valves into alignment, adductor muscles hold valves together
filter feeders, no radula
umbo= oldest part of shell
food particle collected on mucus string in ctenidia to labial palps and into mouth
labial palps surround mouth
foot used for locomotion
siphons are extensions of mantle
most species are semi-sessile in adulthood
-mussels used byssal threads to attach to surface
Phylum Mollusca
Class Cephalopoda
Phylum Mollusca
Class Cephalopoda
-"head-foot"
>squid, octopus, nautilus, cuttlefish
-marine
-largest of all inverts
-smartest of inverts= largest and most complex -central nervous system
-well developed cephalization
-highly developed eyes
-mostly predators= use tentacles to capture food, arms to hold food, beak and radula to tear up food
-closed circulatory system= blood confined in blood vessels, allows for quicker gas exchange, more active lifestyle
-foot modified to form arms, tentacles, funnel (used for expelling water and waste)
-mantle= enclosed visceral mass
-reduce or absent shell, sometimes enclosed
pen= rudimentary shell (squid)
Phylum Annelida
Phylum Annelida
segmented worms
>earthworms, leeches, marine worms
mostly marine, some freshwater and terrestrial
15000 species
Body plan
-organ system
-triploblastic
-eucoelomate
-bilateral symmetry
-cephalized
-segmentation
-protostome
complete gut
coelom functions as hydrostatic skeleton
setae= bristles/hairs, locomotion, holding onto walls of burrows
two-part head
-prostomium= eyes, tentacles, palps
-peristomium= bigger segment around the mouth
Annelida segmentation
metamerism= serial repetition of similar body parts along length of body
-allows for finer control of movement, more efficient movement, protection against injury
-segments= metameres, somites
separated externally (rings= annuli)
internal separation= septa= divide coelom into compartments, composed of peritoneum (epithelial tissue)=lines inner body wall of each compartment, lines organs, septum formed where 2 layers of peritoneum meet
Annelida systems
Nervous
-ventral nerve cord= runs length of body
-fused ganglia in each metamere
-brain= pair of cerebral ganglia
-sensory organs concentrated in head
ocelli, chemoreceptors, statocysts
Respiration
-cutaneous respiration= gas exchange across body wall and cuticle
-a few aquatic taxa have gills
Closed Circulatory system
-may have multiple hearts= aortic arches
Excretory system
-system of paired nephridia in metameres
open to outside by nephridia
collect waste from blood and coelom
Phylum Annelida
Class Polychaeta
Phylum Annelida
Class Polychaeta
-"many hairs"
-marine animals
-includes most annelids
-parapodia= lateral paired appendages, setae covered, locomotion, gas exchange
-acicula= rods that give structure to parapodia, made of chitin
-errant polychaetes
predatious, active, everted jaws on pharynx to capture prey
-sedentary polychaetes
live in burrows or tubes they make in substrate, filter-feeders, don't move a lot
extend tentacles out of burrows or filter out of tube
-reproduction
dioecious
no permanent gonads, temporary swellings that appear during mating season
can sometimes asexually reproduce
Phylum Annelida
Class Oligochaeta
Phylum Annelida
Class Oligochaeta
-"few hairs"= few setae
-mostly terrestrial or freshwater
>includes earthworms
-scavengers
-reproduction
monoecious, hermaphorditic
mutual cross-fertilization
seminal vesicle= where worm makes sperm
seminal receptacles= where worm receives sperm from mating partners
clitellum= reproductive structure (smooth band) made of specialized epidermal cells that secrete muscle
direct development
can also asexually reproduce= fission
Phylum Annelida
Class Hirudinida
Phylum Annelida
Class Hirudinida
>leeches, two smaller groups
mostly freshwater
Hirudinea= leeches
no parapodia, no setae= not haired
somewhat dorsoventrally flattened
dorsal and ventral suckers (front for feeding, back for holding on)
34 segments
-secondary annuli= don't correspond to septa division
fluid feeders
-smaller inverts, blood (sanguivorous)
sanguivorous leeches inject anticoagulant
reproduction
-monoecious
-clitellum only visible during mating season
-similar to oligochaetes
Phylum Arthropoda
Phylum Arthropoda
"jointed foot"
>crustaceans, spiders, insects, etc.
most diverse, successful, widespread, abundant phylum
-1100000 species
marine, freshwater, terrestrial
only flying inverts
carnivores, herbivores, detritivores, omnivores
Body Plan:
-organ system
-triploblastic
-eucoelomate (reduced coelom)
-bilateral symmetry
-well-developed cephalization
-well-developed segmentation
-protostome
Arthropoda strucure
segmentation
-no septa between segments
-segments=somites
-tagmata (tagma)= distinct body regions made up of somites w/ individual functions
exoskeleton= rigid or semi-rigid cuticle secreted by spidermis
-cuticle made of multiple layers that are sclerotized (hardened), made up of chitin and protein and sometimes calcium salts and waxes
-exoskeleton cannot grow with animals= ecdysis
-protection, muscle attachment, shape/structure to body, maintain water balance
-jointed appendages= locomotion, food handling, sensory function, defense or offense
uniramous= one branch end
biramous= two branch end
Arthropoda systems
Nervous
-dorsal brain w/ ventral nerve cord
-ganglia in segments
-sensory organs
ocelli to compound eyes
mechanoreceptors, chemoreceptors, statocysts
auditory organs
complete gut
open circulatory system
-hemocoel= open sinuses where blood can pool
-dorsal heart
hemolymph= blood
reproduction
-most are dioecious
-most do sexual reproduction
-parthenogenesis= virgin birth, some females can produce offspring without male, genetically identical offspring
-direct or indirect development
Phylum Arthropoda
Subphylum Trilobita
Phylum Arthropoda
Subphylum Trilobita
-extinct
-marine, bottom dwellers on substrate
-size: 2 cm to 67 cm long
-scavengers
-dorsoventrally flattened, 3 lobed body plan
-tagmata:
head (anterior), trunk, pygidium (posterior)
-biramous appendagers, 1 pair antennae
legs on all segments except last segment
-mouth parts= hypostome
Phylum Arthropoda
Subphylum Chelicerata
>spiders, ticks, mites, scorpions, horseshoe crabs, sea spiders
-mostly terrestrial, some marine
-tagmata
cephalothorax (head & chest)= legs
abdomen
-uniramous appendages, no antennae, 4 pairs walking legs
-highly modified mouth parts
chelicerae= closest to mouth, first pair of appendages, usually fangs
pedipalps= outside chelicerae, food manipulation, touch
Phylum Arthropoda
Subphylum Chelicerata
Class Merostoma
Phylum Arthropoda
Subphylum Chelicerata
Class Merostoma
>horseshoe crabs
-marine animals
-ancient, only 4 species still extant
-predators of small inverts, nocturnal
-chelicerae and walking legs used to cease prey, pedipalps used as walking legs
-carapace= hard cover of cephalothorax
-ocelli on front, compound eyes on side
-respiration through book gills
Phylum Arthropoda
Subphylum Chelicerata
Class Arachnida
Phylum Arthropoda
Subphylum Chelicerata
Class Arachnida
>spiders, scorpions, ticks, mites, harvestmen
-terrestrial
-respiration through book lungs
-mostly free living, some parasites
-mostly predators
-chelicerae and pedipalps modified
spiders= chelicerae have fangs to inject venom, pedipalps have sensory function, males use pedipalps for sperm transfer
scorpions= chelicerae for shredding food, pedipalps modified into claws for ceasing prey
Phylum Arthropoda
Subphylum Myriapoda
>centipedes, millipedes
-lots of parts
-all terrestrial
-tagmata:
head, trunk (repeated segments)
-uniramous appendages, 1 pair antennae, 5+ pairs walking legs
-mouth parts= paired mandibles for chewing
-respiration through system of tracheae= tracheal system
system of tubes throughout body
open to outside through holes= spiracles
Phylum Arthropoda
Subphylum Myriapoda
Class Chilopoda
Phylum Arthropoda
Subphylum Myriapoda
Class Chilopoda
>centipedes
-predators= active, fast moving
-venomous
-dorsoventrally flattened
-varying number of segments, depends on species
-1 pair spiracles per segment= one on each side
-1 pair legs per segment
first segment has venom claws instead of legs, last 2 segments lack leggs
Phylum Arthropoda
Subphylum Myriapoda
Class Diplopoda
Phylum Arthropoda
Subphylum Myriapoda
Class Diplopoda
>millipedes
-detritovores, herbivores
-slow moving, coil up for protection
-secrete toxins out of side of body
repugnatorial glands= where toxins secreted out of
-round, cylindrical body
-varying number of segments
-2 pairs spiracles & 2 pairs walking legs per segment
Phylum Arthropoda
Subphylum Crustacea
Phylum Arthropoda
Subphylum Crustacea
second most speciose
crusta= "shell"
mostly marine, some terrestrial & freshwater
tagmata:
-cephalothorax, abdomen
biramous appendages, often 5 pairs walking legs, 2 pairs antennae, 2 paired chewing madibles
modified appendages
-mouth parts
antennules, antennae= sensory function
maxillae= hold food
maxillipeds= holding/manipulating food
mandibles= chewing
-chelipeds= grasping food, defense (big claws on end= chela)
-walking legs= walking, crawling
-swimmerets= swimming
males: copulatory swimmerets= sperm transfer
-uropods= part of tail, locomotion, direction
female: used to protect eggs
respiration= gills
-smaller crustaceans= gas exchange across thinner areas of cuticle
excretory system= antennal gland/green gland, gills, or across thinner areas of cuticle
-antennal gland=excretory functions, water balance
Phylum Arthropoda
Subphylum Hexapoda
-six leg animals
-mostly terrestrial, some freshwater
-tagmata
head, thorax, abdomen
-uniramous appendages, 1 pair antennae, 3 pairs walking legs, mandibles for chewing
Phylum Arthropoda
Subphylum Hexapoda
Class Insecta
Phylum Arthropoda
Subphylum Hexapoda
Class Insecta
ecologically, economically, medically important
found almost anywhere except in oceans
free-living & parasitic
eat everything
small size, wings, high adaptability, no competition (intraspecific competition) between larvae and adults= allow for wide distribution
wings
-2 pairs as adults (larvae have none), not appendages
-true flies have 1 pair, second pair modified into haltere= functions as gyroscope= balance, orientation
-some species have lost wings entirely (fleas, lice)
excretory system
-malpighian tubules=set of tiny tubes w/ ends in hemocoeal to filter out wastes
respiration= tracheal system
reproduction
-dioecious
-sexual reproduction
-metamorphasis= extreme morphological changes from egg to adult
instars= developmental stage between molts
(2-20 instars, depends on species, environment)
Phylum Echinodermata
Phylum Echinodermata
"spiney skin"
>sea stars, sea urchins, sand dollars, etc.
all marine
ancient and highly derived
Body plan:
-organ system
-triploblastic
-eocoelomate
-pentaradial symmetry (larvae= bilateral)
-no cephalization
-no segmentation
-deuterostome (anus forms fist)
complete gut
open circulatory system
most closely related to chordata
oral-aboral axis (oral=bottom, aboral=top)
no brain, decentralized nervous system
-nerve ring w/ nerves radiating out into each arm
-not well developed sensory receptors
reproduction
-dioecious
-indirect development
larval stage has bilateral symmetry
Echinodermata structure
Water-vascular system
-locomotion, gathering food, excretion, gas exchange
-system of canals filled with sea water throughout animal
-madreporite= large pore in which water enters
-stone canal
-ring canal= circles central disk
-radial canals= radiate from each arm
-lateral canals= extensions off of radial canals, connect rest of system to tube feet
-ampulla= sacs attached to tube feet, regulates water pressure in tube foot
ampulla filled with water can contract and fill tube feet for use

Endoskeleton
-calcareous plates= dermal ossicles=plates made of calcium carbonate, embedded in skin, covered in epidermis
-movable spines= defense, locomotion
-pedicellaria= defense, protrude through space between dermal ossicles, pincers, remove debris from aboral surface
-dermal branchia= protrude through space between dermal ossicles, skin gills, used for respiration
-tube feet= locomotion, respiration
ambulacra=grooves in which tube feet are found
Phylum Echinodermata
Class Asteroidea
Phylum Echinodermata
Class Asteroidea
>sea stars, sun stars
-5 or more arms connected to central disk
-mouth on oral surface to substrate, anus at top
-dermal ossicles= loosely bound, gaps between plates
-predators, scavengers, will eat any slow-moving invert
bivalves, sea urchins
-regeneration= if looses arm, it can grow a new one
lost arm can grow new sea star if enough of disk remains
Phylum Echinodermata
Class Crinoidea
Phylum Echinodermata
Class Crinoidea
>sea lilies, feather stars
-sessile= stock w/ attachment to substrate, occasionally motile
-filter feeders
-5 arms with branches
-upper surface= oral surface
mouth and anus on oral surface
-water vascular system has coelomic fluid instead of sea water
-dermal ossicles articulate with each other, closer together
Phylum Echinodermata
Class Ophiuroidea
Phylum Echinodermata
Class Ophiuroidea
>brittle stars, serpent stars, basket stars
-"serpent tailed"
-predators, scavengers, filter-feeders
-5 arms= thinner and distinct from central disk
use arms for locomotion instead of tube feet
-oral surface faces substrate (down)
mouth and madreporite on oral surface, no anus
-dermal ossicles articulate closely with each other
-regeneration= can cast off arm to escape predator
-reproduction= sexual (asexual= can cleave disk in half)
Phylum Echinodermata
Class Echinoidea
Phylum Echinodermata
Class Echinoidea
>sea urchins, sand dollars, heart urchins
-"spiney"
-herbivores, some scavengers
often preyed on by other echinoderms
-fused or tightly joined dermal ossicles= form a shell= test (under thin layer of epidermis)
-no arms, still pentamerous plan
-Aristotle's lantern= 5 part jaw structure
Phylum Echinodermata
Class Holothuroidea
Phylum Echinodermata
Class Holothuroidea
>sea cucumbers
-mostly filter feeders, few detritovores
-reduced dermal ossicles
-elongated oral-aboral axis= parallel to substrate
mound on oral surface with tentacles (modified tube feet)
anus on aboral surface
-sole= surface that rests on substrate
3 ambulacra on sole, 2 on top
-autonomous evisceration
when disturbed, they go through strong muscular contractions the expel guts, used to escape predators