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81 Cards in this Set

  • Front
  • Back

What are species?

Groups of organisms that interbreed and share a suite of genetic and morphological attributes, and are reproductively isolated from other such groups.

What is speciation?

The divergence of biological lineages and the emergence of reproductive isolation between lineages, creating separate species.

What are the three major classes of species concepts?

Morphological, biological, and lineage.

What does the morphological species concept state?

A species comprises individuals that look alike and that individuals that do not look alike belong to different species.

Why does the morphological species concept work?

Members of the same species share alleles that code for the same morphological features.

What are the two limitations of the morphological species concept?

1. Members of the same species do not always look alike, like the males, females, and young individuals.



2. Cryptic species, which look exactly the same and indistinguishable, do not interbreed.

What does the biological species concept state?

Species are groups of actually or potentially interbreeding natural populations which are reproductively isolated from other such groups.

What is reproductive isolation?

A state in which two species can no longer exchange genes.

What are the three limitations of the biological species concept?

1. Does not work with closely related individuals.



2. Does not apply to organisms that reproduce asexually.



3. Limited to a single point in evolutionary time.

What does the lineage species concept state?

Species split into two descendant species which thereafter evolve as distinct individuals and are considered over evolutionary time.

What is a lineage?

An ancestor-descendant series of populations over time.

What is the most important idea for the origin of species?

Reproductive isolation

What is the Dobzhonsky-Muller Model?

A mode that explains how reproductive isolation results from accumulation of genetic incompatibilities following a splitting of an ancestral population, interrupting gene flow.

By the Dobzhonsky-Muller Model, what would happen to the offspring of the two genetically incompatible individuals (looking at alleles)?

The hybrid offsprings alleles would be either functionally inferior or lethal causing death.

T/F:


Greater geneteic differences and incompatibilities lead to increased reproductive isolation, and thus speciation.

True

What is allopatric speciation?

Speciation resulting when a population is divided by a physical barrier.

What is the dominant and most common mode of speciation?

Allopatric speciation

Whta is sympatric speciation?

Speciation without a physical barrier causing reproductive isolation.

Whate are the two ways sympatric speciation occurs?

1. Disruptive Selection where individuals have preference for distinct microhabitats where mating takes place.




2. Polyploidy where there is a duplication of sets of chromosomes.

What is the most common way sympatric speciation occurs?

Polyploidy

What is autoploidy?

Chromosome duplication in a single species, so internal.

What is alloploidy?

Chromosomes combining from two species, so external kinda.

How does autoploidy occur accidentally?

If two diploid genes combine, resulting in a tetraploid individual. When the tetraploid and diploid mate, a triploid is made that is sterile. Therefore, tetraploid can only self-fertilize or mate with another teraploid.

Why is polyploidy important in plant evolution?

70% of flowering species and 95% of fern species are the result of polyploidization because many plants can self-fertilize.

What are reinforcement mechanisms?

If hybrids are less fit than non hybrids, selection will favor parents that do not produce hybrid offspring. This reinforcement prevents hybridization.

What are prezygotic isolating mechanisms?

Mechanisms that prevent hybridization from occuring before fertilization.

What are postzygotic isolating mechanisms?

Mechanisms that reduce fitness of hybrid offspring, selecting against hybrizization.

What are the five prezygotic isolating mechanisms?

1. Mechanical


2. Behavioral


3. Gametic


4. Temporal


5. Habitat

What is mechanical isolation an what is an example?

Isolation caused by differences in size and shape of reproductive organs that make mating impossible. For example, orchids pollinators.

What is temporal isolation and what is an example?

Isolation where mating periods do not develop because the species have different mating seasons. For example, 3 closely related frogs that live in the same habitat breed at different times of year.

What is behavioral isolation and what is an example?

Isolation due to the fact that individuals reject or fail to recognize potential mating partners. For example, breeding calls of male frogs quickly diverge between related species, so the female frogs ignore calls from other species.

What is habitat isolation?

Isolation where when two closely related species evolve preferences for living or mating in different habitats, causing them to never come into contact during mating periods.

What is gametic isolation?

isolation where th eggs of one species do not have appropriate chemical signals for sperm of another species; or sperm is not able to attach to and penetrate the egg.

What are the 3 postzygotic isolating mechanisms?

1. Low hybrid zygote viability where hybrid zygotes fail to mature normally or have severe abnormalities causing death.


2. Low hybrid adult viability where hybrid offspring may have lower survivorship than nonhybrids.


3. Hybrid infertility where hybrids may mature into infertile adults.

What are hybrid zones?

When, unless reproductive isolation is complete, closely related species may hybridize in areas where their ranges overlap.

What are the 4 factors that can influence rates of speciation across groups?

1. Diet specialization


2. Pollination


3. Sexual selection


4. Dispersal ability

What is diet specialization and what is an example?

When populations with specialized diets are more likely to diverge. For example, herbivorous bugs specialize on one or a few plant species.

T/F:


Herbivorous beetle groups contain many more species than the related predatory species do.

True

What does pollination do for speciation?

Speciation rates are higher in animal-pollinated than wind-pollinated plants.

What does sexual selection do for speciation and what is an example?

The mechanisms of sexual selection result in high rates of speciation. For example, the birds of paradise in New guinea.

How does dispersal ability affect speciation and what is an example?

Speciation rates are likely to be faster in species with poor disperal abilities because they can be separated by even narrow barriers. For example, land snails with many species may be restricted to a single valley.

Rapid speciation can lead to evolutionary radiation. What is evolutionary radiation?

Proliferation of many species from a single ancestor.

What is adaptive radiation and what is an example?

If the resulting species from evolutionary radiation live in a wide array of environments. For example, the Haiwaiian Islands and the 10,000 native species of insects that are believed to have evolved from 400 immigrant species.

What is a genome?

An organisms full set of genes plus noncoding regions of DNA.

T/F:


In Eukaryotes, some DNA is in the mitochondria and chloroplasts.

True

What is molecular evolution?

The process of change in the sequence composition of cellular molecules such as DNA, RNA, and proteins across generations.

What 3 ideas does molecular evolution include?

1. Mechanisms and consequences of macromolecule evolution.


2. Relationships between gene structure and proteins and organism function.


3. using molecular variation to reconstruct evolutionary history.

What does homologous mean?

Shared by two species or inherited from a common ancestor.

What is sequence alignment and its technique?

Aligning amino acid sequences and determing insertion and deletion events that have occured since the two species diverged.

What is a similarity matrix?

A computer algorithm matrix that gives a measure of the minimum number of changes that have occured since the divergence of each pair of organisms.

What is a limitation of a sequence alignment?

The count provided of the changes in DNA underestimates the actual number of changes that have occurred since the sequences diverged.

What are multiple substitutions?

More than one change has occurred at a given position between the ancestral sequence and at least one of the observed sequences.

What are coincident substitutions?

At a given position, different substitutions have occurred between the ancestral sequence and each observed sequence. Basically, different substitutions in different descendants.

What are parallel substitutions?

The same substitution has occurred independently between the ancestral sequence and each observed sequence.

What are back substitutions?

Also known as reversions. After a change at a given position, a subsequent substitution has changed the position back to the ancestral state.

What is a transversion change?

Purines are replaced by a pyrimidine and vice versa.

What is a transition change?

Changes between the two purines A to G, or changes between pyrimidines C to T and vice verse. More common than transversions.

What is the start codon for methylene?

AUG

What are some stop codons?

UAA, UAG, and UGA

What is a synonymous or silent mutation?

Substitutions that do not change the encoded amino acid.

What is are the two nonsynonymous substitutions?

Misense substitutions and nonsense substitutions.

What are misense substitutions?

Changes in a specified amino acid.

What are nonsense substitutions?

Produces a stop codon and terminate the protein.

What is positive selection in terms of substitutions?

A replacement that confers an advantage to the organism.

T/F:


Nonsynonymous substitutions are likely to be deleterious but may be selectively neutral if protein shape and function are not altered.

True

What are pseudogenes?

Copies of genes that are no longer functional (and so not subject to strong selection). Has a much higher rate of substitution than regular genes.

What does the neutral theory state?

The majority of mutations are selectively neutral and accumulate through genetic drift, not positive selection.

How are molecular clocks used?

Used to calculate evolutionary divergence times between species if mutation rates are constant.

T/F:


There is some correlation between genome size and organism complexity but larger does not always equal more complex.

True

What are 3 possible roles of non coding DNA?

1. Regulatory functioncontrolling timing of gene expression.


2. Pseudogenes that are raw material for evolution.


3. Transposable elements that reproduce faster than the host genome.

What are retrotranspons?

Transposable elements that copy themselves through an RNA intermediate. Commonly carry long terminal repeats.

T/F:


The number of orphaned LTRs (long terminal repeats) equals the number of retrotranspons that have been lost.

True

What is lateral gene transfer?

Genes, organelles, or gene fragments moving horizontally from one lineage to another, seen commonly in plants.

What are two advantages to lateral gene transfer?

1. genes that have antibioticresistance are transmitted among bacteria species.


2. Increases genetic variation, providing new material for evolution.

T/F:


Lateral gene transfer is relatively uncommon among most eukaryote lineages but endosymbiosis can be viewed as lateral transfer technically.

True

What are 4 possible fates for when gene duplications give rise to new protein functions?

1. Both copies retain original function with more product created.


2. Both copies retain original function but expression diverges in different tissues or at different times.


3. Once copy accumulates deleterious substitutions and becomes either pseudogenes or eliminated.


4. Once copy accumulates substitutions that allow it to perform a new function.

What are gene families?

Genes created from successive rounds of duplication and mutation, like the globin gene family.

T/F:


In polyploidy organisms, the whole genome is duplicated, allowing new functions to arise like with jawed fish.

True

What are the two forms of homology in genes since members of gene families have similar sequences?

1. Orthologs


2. Paralogs

What are orthologs?

Homologous genes traced back to speciation events (external).

What are paralogs?

Homologous genes related through duplication events (internal).