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Phylum Nemertea- What are they?

Ribbon or Proboscis Worms

What is another name for them?

T/ F:


Phylum Ectoprocta are Triploblasts

True

Phylum Ectoprocta

-"goblet worms"-characterized by solid tentacles that form a circle- look like hydroid cnidarians, have ciliated tentacles.

Tiny phylum- look like mini goblets

Phylum Cycliophora

-very small (size and species number)-Only on the mouth of certain lobsters

Lobsters

Lophotrochozoa

Belong to bilateria, are protostomes.


-Include: Byrozoa, Platyhelminthes, Annelida,Brachiopoda, nemertini, mollusca,

Phylum Nemertea

-1300 species (terrestrial and marine)


-Known for Proboscis in Rhynchocoel cavity (mesodermal=coelom)


- uses their esophogus as a muscle to grab its food.


-proboscis can be slimy or poisonous

Remember weird worm and videa

Trochozoans

-small planktonic larva with rings of cilia around the mouth


-uses cilia for swimming and getting food

Phylum Mollusca

-Super diverse (85,000 species)


-Disparate (8 classes!)


- most are small but also include giant squid!

2nd to Anthropods in species

Molluscs are important to us?

1. Food (we eat lots of them)


2. We use pearls and shells as currency


3. Bioindicators- can show us when environment is getting bad, and filter water (like oysters)


4. pests- can be invasive (zebra mussels), intermediate hosts


5. Research- big nerve cells and giant axons, also strongest biological structure on earth- abalone shells)

5 Reasons!


FCBPR

What are 2 characteristics of a mollusc?

1.Visceral Mass


2. Head foot

Head foot includes:

-needed for sensory and locomotion


-most have a well developed head with sense organs (brain, eyespots, tentacles)





what is it needed for?

Radula

-Not FOUND IN BIVALVE


-Chitinous ribbon of teeth, that moves over an odontophore cartilage and can odontophore can move it in and out.

https://www.youtube.com/watch?v=cEMK3VN8pW8

The Foot

-used for locomotion (cilia, muscular contractions, and mucus),


Cephalopods, modified siphon for jet propulsion

Visceral Mass

-contains the internal organs, (digestive, excretory, respiratory, and reproductive)


-includes the mantle and mantle cavity

Mantle

-"the skin"


-can have muscles or chemoreceptors


-makes the shell if the organism has one


-the surrounding tissues

Mantal Cavity

Open to "world"


-where the folds connect


-can be for excretion


-where it poops, pees, reproduction, and O2 exchange

Three layers of Mantle

(made by outer layer of mantle)


1. Periostracum


2.Prismatic Layer


3. Nacre

Periostracum

Outer organic layer (resistant protein conchiolin)


-is like the skin of a shell



(outside)

Prismatic Layer

-Thickest layer


-calcium carbonate plates in proteinacious mix (some of the strongest material on earth)

-super strong!

Nacre

"pearly layer"


-pretty, shiny, smooth layer


-continuously thickens


-where pearls come from!


-calcium carbonate and protein sheets

-think of pearls

Explain Pearls!

-form when irritating debris gets between shell and mantle


-innermost layer is nacreous layer, which continously grows


-coats in layer of nacre

Internal characteristics of Molluscs

-Bilaterally symmetric coelomates (reduced coeloms)


-HEART IS ONLY COELOMATIC CAVITY


-Mantle/mantle cavity used for respiration


-Most have open circulatory system (cephlopods have closed)


-Complex digestive system


-Nervous system varies, large nerve cells

Reproduction is Molluscs

-Always sexual


-diocieous or monoecious


- some have TROCHOPHORE larvae (aquatic larvae)


-Veliger larvae common in aquatic forms- mini mollusc (has an early shell and ciliated foot like structure)


-Cephlapods and Land snails have direct development

Molluscs


Classes Caudofoveata and Solenogastres (The Aplocophorans)

-wormlike and shell-less


-calcareous sclaes/spicules


-marine detrital/microorganism consumers (burrowers)


-reduced head, no foot

Molluscs


Class Monoplacophora

-"one plate"


-thought to be extinct


-round shell, large foot


-serial repetition of body parts


(gillsm nerves, gonads, nephridia)

-resemble limpets in the gastropods

Mollusc


Class Polyplacophora

-"many plates"- kind of like armadillos, have 8 movable plates


-Mantle girdle around outside


- Serial repetition seen, lots of internal and extermal structures like gills and nerves


-Found in the intertidal rocky areas

Molluscs


Class Scaphopoda

-"tusk shells/tooth shells"


- Sedintary


-tubular shell is open at both ends, stick and called the CAPTACULA


-tentacular foot for burrowing and food capture


-Oxygen diffuses right through internal walls in the mantle

Mollusc


Class Gastropoda

"stomach foot"


-most diverse class (60,000+)


-includes snails, limpets, sea slugs, periwrinkles


-Marine, freshwater, terrestrial


-shell is the main defence (because they are typically pretty slow)

Think snails

3 main groups of gastropods

1. Prosobranchia


-most marine smails (limpets, conchs)


2. Opithobranchia


-mostly marine, shell-less forms and have no gills- instead they have dorsal extensions called CERRATA. (POISONOUS)


3.Pulmonata


-most land and freshwater snails and slugs (mantle cavity into lungs)



Torsion

-180 degrees rotation of the mantle/mantle cavity


-happens in VELIGER larval stage

Coiling

Whorling of the shell to the right side of the body (results in the spiral on the shell), no right mantle cavity.


-also happens in Veliger larval stage, but evolved first)

Feeding in gastropods

-all feed with some variation of a radula


-some scrape algae, eat dead fish and drill into other mollusc shells and suck out their insides.


-herbivorous, carnivours, scavengers


-cone snail- proboscis that harpoons fish

Gastropod SEX

-can be dioecious or monoecious


-fertilization is internal


-Marine forms VELIGER LARVAL stage


-terrestrial, baby snails


-Simultaneous hermaphrodites, share and ACCEPT sperm.


-both accept sperm, but stab eachother in the head to put it there.


-love darts are mating sperm booster


-have orgies



Mollusc


Class Bivalvia- "two valves"

-no head, no radula


-laterally compressed shell


-hinged dorsally, gape open ventrally


-sedentary suspension feeders use mucous covered gills, water flow created by its beating cilia.


-burrowing forms(clams) have a large musclular foot from the anterior/ventral end.


-siphons are made from the mantle


-umbo is oldest part of the shell (hinge)


-Adductor muscles open and close the shell

Bivalve internal features

Gills- (called CTENIDIA) used for feeding and respiration


- good food is sorted into bolus in the anterior end where it gets eaten

Bivalve locomotion

-are mostly filter or suspension feeders using CTENIDIA.


-use foot to burrow


-adductor muscles to swim (scallops to swim)


-And some are sessile,


-Byssal threads-(mussels)


-Oysters secrete a cement to attach themselves to rocks and such

Cephlopoda

-"the head foot"- indistingishable


(siphon comes from the foot)


-exclusively marine




-4 distinctions:


-squids


-octopuses


-cuddlefish


-Nautiluses




Shells in Cephlopods

Nautiloids- large shells (remnants of ancient group AMMONITES) shell gas regulates!


Cuttlefish -internal shells in mantle


Squid- Proteinacious pen


Octopus- no shell at all

Locomotion in Cephlopods

- fins, arms, and jet propulsion (shoots water out of the siphon-modified foot) to shoot the animal backwards.

Basic Body Plan of Cephlopods

- 1 pair of gills


-8 arms (squid and cuttlefish have extra 2 tentacles)


-CLOSED circulatory system w/ multiple hearts


-Nervous system includes big brain, large lensed eyes and great control over arms

Camouflage and Ink in Cephlopods

-4 cell types (chromatophores)


- Ink sac- melanin and mucus, comes out the rectum.

Reproduction in Cephlopods

Dioecious


-lots of competition for mates


-sperm is passed in packets to female


-eggs are laid on sea floor, and are watched by females


-direct development, out of egg pops baby cephalopod

ANNELIDS as Lophotochozoans

-protostomes


-coelomates


-triploblastic


-trochozoans



COMPARE ANNELIDS TO MOLLUSCS

-Both are greater lophotrophozoans


-Annelids coelom allows for hydrostatic skeleton


- Annelids can change shape rapidly




Molluscs- shell prevents large shape changes


-Coelom reduced to pericardial cavity

Annelid locomotion

-Coelom filled with fluid, creating hydrostatic skeleton


-contraction of longitudinal muscles- body shortens and expands


-contraction of circular muscles, body narrows and lengthens


-Peristalsis- alternating contractions


-swimming annalids use undulatory movements (muscles)

Phylum Annelida- "ringed worms"

-also called "bristle worms"


-Setae (chiton little bristles)


-15,000 species (marine, freshwat, damp terrestr.)


-all lifestyles, deposit feeders, predactors, suspension feeders, parasites (leeches).


-closed circulatory system with pumping hearts


-centralized nervous system w/ good cephalization

Defining Annelid Features

-Setae- chitinous, epidermal bristles


-helps in burrowing, like little anchers




Parapodia- "paddle-like feet"


-used in swimming and respiration (used like gills)





Annalid basic Body Plan

-2 part head (prostomium and peristomium)


-numerous segments


-Terminal Pygidium (kinda like the butt): new segments


-epidermis coverented in thin-non chitinous cuticle

Pleistoannelida: Errantia

-motile poluchaetes "many long hairs"


-mostly marine


-well developed sense organs and cephilization


-ARE POLYCHAETES

Pleistoannelida: sedentaria

- contains former olichaetes, echira, etc.

Sedentaria Polychaetes

-cilia and mucus and food groves in tentacles

Sedentaria: Siboglinidae

"beared worms"


-eat via absorption, deep sea, chill near hydrothermal bents or endosymbiotic bacteria

Sedentaria: Echiuridae

-unsegmented, flattened

Sedentaria: Clitellata

-3000 species


-earthworms!


-well developed circulatory, digestive, and excretory system!


NEPHRIDIA- cool little funnel that filters coelomic fluid

Carnivorous Leeches

Kinabalu giant red leech from Borneo- only eats other worms, about 30 cm long

Phylum SIPUNCULA

-the Peanut worm


-250 species


-not segmented and no setae


-ciliated introverted feeding tentacles


-not annelids but they are their closest relative!

KNOW THIS STUFF!!!

RECAP OF ANNELIDA

(according to new tb)


-Errantia and Sedentaria


-errantia- motile polychaetes


-sedentaria= tube/burrow polychaetes, siboglinidae, Echiuridae, and Clitellata (contains Oligochates and Hirudinida)


-Phylum Sipunula

Sedentaria: Clitellata

- formally Oligochaeta and Hirudinida classes


-no Parapodia


-all hermaphrodites


-have direct development (no trochophore)

Ecdysozoa

"the molting protostomes"


-Good monophyletic group


-everyone in it can molt

Ecdysozoans are : (3 key transitions)

Protostomes, triploblastic, and bilaterally symmetic!


-no locomotory cilia


-hard overing exoskeleton made of collagen or chitin

PHYLUM NEMATODA

Up to 80% of all life on earth!


-can be freeliving or parasitic


-Pseudocoeulomates


-collagenous cuticle

Nematoda body plan

-Tube within a tube


-muscular pharynx sucks in food


-good reproductive system- diocous

First animal to have its genome sequenced?

C. Elegans (Nematode)

Nematoda as parasites

-cuticle=defence


-adapted mouth parts (hooks and burrowing)


-reproduce lots for egg to infect organism


-Roundworms (can block your intestines)


-Hookworms (burrow into skin)


-Filarial worms (on anthrodods), infect lymph nodes

Phylum Nematomorpha

-horsehair worms


-can grow up to 1m


-semiparasitic (adults are freeliving but juveniles live in arthropods

Phylum Onychophora

-"velvet worms"


-soft cuticle- hydrophobic (looks like velvet hence the name)


-all predators of insects

Phylum Tardigrada

-"water bears"


-sucking pharynx- eats plant and animal via juices


-Cryptobiosis!!!

Cryptobiosis

-called a Tun in seabears


-go into ametabolic state, pretty much dead until favourable conditions apply


-can survive almost anything!!!

Phylum Anthropoda!

= joint foot


-80% of named animals


-segmented bodies


-joint limbs


-chiton exoskeleton


-

Anthropod Key transitions!

-Organ system


-bilateral symmetry


-coelomate


-triploblast protostome


-yes to segmentation

Exoskeleton in Anthropoda

Chitinous cuticle; used for protection and water retention.


-lightweight so doesnt get in the way


-when thin, it can be permeable

Segmentation and appendages in Anthropoda

-more effienct movement


-specific segments have specific functions

Respiration in Anthropoda

-some breathe air, some water


-efficient for fast metabolism

Sense organs in Anthropoda

-eyes


-hearing


-balance


-chemosensory

behavioural adaptations in anthropoda

-true sociality


-think of leafcutter ant

metamorphosis

-lots of niches for the same species


-larvae eats something different than adult


-more environments, more numbers!

Exoskeleton, segments, and appendages

-group of segments called Tagmata (work together for common function)


-"head, thorax, and abdomen"

Subphylum Chelicerata

-horseshoe crap, spiders, scorpions


-TWO tagmata (cephalothorax and abdomen)


-Appendages: Chelicarae and Pedipalps with 8 legs


-NO MANIBILES OR ANTENNAE

Subphylum Myriapoda

-like centipedes


- 1 pair of antennae


-mandibles


-centipedes and Millipedes


-Head and (LOOOONG)Trunk

Subphylum Crustacea

-mostly marine


-two pairs of antennas


-two pairs of maxillae


-Biramous- means two branched- endopod, exopod

Crustacean body plant

-Carapace: chiton exoskeleton, covers head and thorax.




-Have a HEAD, THORAX, ABDOMEN

Crustaceans internal form

-open circulatory system


-Gills attached to appendages


-ANTENNAL glands take care of waste!

Crustaceans Sensory system

-advanced nervous system


-developed eyes, and taste and smell from chemosensory

Reproduction in Crustaceans

-most are diocious


-brood eggs in brood chambers


-have larval stage!!!

Feeding in Crustacea

Maxillae and Mandables- think claws that shred food.


-claws and walking legs can also capture food

Ecological importance of Crustacea

-are zooplankton, bottom of the ocean food chain


-mutualistic relationships with other critters


-parasite hosts!


-huge fishery for them!!!

EXAMPLES OF CRUSTACEA:


Ostracods

-bivalve and resuced thoratic appenadages

EXAMPLES OF CRUSTACEA:


Branchiopods

-lots of freshwater species, brine shrimp

EXAMPLES OF CRUSTACEA: Copepods

Seen throughtout the entire life, no legs or carapace

EXAMPLES OF CRUSTACEA: Barnacles

sessile as an adult, no sephilization

Largest Penis to body ration

Barnacles!

Malacostraca Crustaceans

-Isopods- pill bugs


Euphasiacea- KRILL


Decapods, 10 legs like lobster! (first legs are claws)

SUBPHYLUM HEXAPODA

-6 legs (DUH)


-all uniamous (the same)


- three tagmata (HEAD, THORAX, ABDOMEN)

Class Insecta

-found everywhere, exoskeleton is light and keeps water in :)

Characteristics of Insects

-Head, thorax abdomen


-3 pairs of legs and 2 pairs of wings on THORAX


-have MANDABLES!

Only Arthopod without Mandables

Chelicerates!

WIngs

-NOT APPENDAGES, simple extensions of cuticle on thorax


-usually 2 pairs


-only invertebrates to fly


-Direct flight- muscles attache to wings


Indirect- alter the shape of thorax for wing movement


-moves in a figure 8

Insect respiration

-tubular heart, no blood for oxygen, just nutrients


-spiracles have opening trachae- gas exchange over wet membrane

Insect reproduction

-dioecious


-internal fertilization


-female lays eggs after fertilization

Insect developement

- Molt between stages


-HOMOMETABOLOUS- complete (egg, larva, pupa, adult- butterfly)


-HEMIMETABOLOUS (incomplete), egg, nymphs, adult

Insect social behanviour

-Eusociality- class system


-some workers, drones, and then a queen


-think of bees/ants!

DEUTEROSTOMIA

-Radial cleavage


-regulative (if split, makes whole new embryo)


-Blastopore makes anus- mouth comes second


-Entercoelous- coelom forms by outpocketing the mesoderm.

5 transitions Echinodermata

-Organ system


-Pentaradial symmetry


-Coelomate


-triploblastic & DEUTOERSTOME


- NOT segmented

Phylum Echinodermata


(STARFISH)

-spinky skin


-marine


-no cephilization



Phylum Echinodermata: exoskeleton

- part of special coelomic cavity called stereom, made of collegen

Water Vascular System

Hydraulic system


-canals and tube feet


FUCTION- respiration, excretion, locomotion and feeding

SERIES OF CANALS

MADREPORATE, STOME CANAL, RING CANAL, RADIAL CANAL, AMPULLAE, TUBE FEET




MSRR CAT

Pedicellariae

-on aboral surface


-clean, foodcapture, defence

Starfish respiration

-Papulae (skin gills)


-increase surfae area for respiration

Ambulacra-

band from which tube feet form

Internal characteristics of Starfish

-no brain, epidermal nerve net thought


-digestive system varies!


-often seperate sexes with EXTERNAL fertilization

Larval Devolopement in starfish

-start as bilateral, remember they are DEUTEROSTOMES, larvae gets early tube feet!

Starfish regeneration

-just need 1/5 central disc and 1 arm


- Autotomy: deliberate shredding of body parts or appendages to escape

Fragmentations in Starfish

-form of assexual reproduction


-"comets"


-just arm needed

ECHINODERM CLASSES:


Crinoidea

-"lily form"


-oral side up


-NO PEDICELLARIEA, MADREPORATE, or SPINES

ECHINODERM CLASSES:


Asteroidea

-common sea star


-central disc and arms


-2 part somach- everts tomach


-carvivours





KETSTONE ECHINODERMS

-PURPLE INTERTIDAL seastar controls mussel populations-CROWN OF THORMS, kills coral reefs

-Pacific Sunflower- eats everything

Class Ophiuroidea-

-brittle stars


-really good at regeneration


-all organs in ventral disc


-

Class Echinoidea

-Sea urchens and sand dollars


-compact body form


-mouth is called ARISTOTLES LANTERN


-compact bodies- Ossicles as plates

Urchin Barren

-kill the kelp, so sea otters control the urchin population!

Class Holothuroidea

-Sea cucumbers


-elongated on oral-aboral axis


-reduced occicles


-ventral tube feet for locomotion


-oral tentables are modified tube feet

Sea cucumbers

-have a big coelomatic cavity


-Cloaca- "all purpose orifice


- 1 true Pelagic species

PHYLUM HEMICHORDATA

closely related to echinoderms


-have a "notochord" called a stomochord"


-not actually half chordate, just thought to be and never changed the name.