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19 Cards in this Set

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Define:

Speciation
The formation of a new species from a pre-existing species

Occurs at the branching events in a phylogeny (cladogram)
What are the 4 main modes of speciation?
1) Allopatric

2) Parapatric

3) Sympatric

4) Hybridization
Define:

Allopatric speciation
Populations become separated by a physical barrier and then evolve reproductive barriers.

Outlined by Ernst Mayr.

*The most common* mode of speciation.

Two types:
- Vicariant speciation
- Peripatric speciation
What is the most common of the 4 modes of speciation?
ALLOPATRIC
Define:

Vicariant speciation
A form of allopatric speciation.

Population is divided into two widespread populations by a physical barrier
- e.g. changes in courses of rivers, mountain-building events, Volcanism, glaciation, environmental changes
What are some different types of barriers that can lead to vicariant speciation?
Changes in courses of rivers

Mountain-building events

Volcanism

Glaciation

Environmental changes
What were 2 examples given for vicariant speciation?
1) Speciation in 6 pairs of freshwater fish in southeastern U.S.
- split into a western and eastern clade.

2) Snapping shrimp
- Separated on either side of the Isthmus of Panama
Define:

Peripatric speciation
A type of allopatric speciation.

A colony disperses from a widespread population to a new area and evolves reproductive isolation.
What plays a large role in accelerating the process of peripatric speciation (once the initial move happens)
Genetic Drift

The new population is small, thus genetic drift will have a large influence.
What organism (and geographic location) was used as an example for Peripatric Speciation?
Hawaiian fruit flys across the Hawaiian Islands.
Summarize the characteristics of Vicariant Speciation (3 points) and Peripatric Speciation (3 points)
Vicariant Speciation
- Physical barrier
- No dispersal
- Results in 2 large populations


Peripatric speciation:
- Physical barrier
- Dispersal
- Results in 1 large and 1 small population
Define:

Parapatric speciation
Strong selection causes a continuous population to diverge

Populations are contiguous (boundaries are in contact)

No physical barrier present

Divergent selection may be strong enough to overcome gene flow; reproductive isolation may evolve.
What organism was used as an example for Parapatric Speciation?

What caused it?
Grass species: Anthoxanthum odoratum

Evolved different flowering times due to natural selection from mine wastes.

Evolution of reproductive isolation over a short distance.
Define:

Sympatric Speciation
Evolution of reproductive isolation within a randomly mating population

No geographic separation.
What organismz were used as examples for sympatric speciation?
Pyrenestes - Seedcracker Finches.

Insect host switching trees (Apply fly maggot)
Define:

Hybridization Speciation
Two parental species hybridize and produce offspring that can breed with each other, but are reproductively isolated from the parent species.

*Important in plants to get new species. Rare in animals.

Chromosomal barriers may isolate offspring from the parents.
Define:

Speciation by polyploidy
A type of hybridization speciation.

- Errors in mitosis or meiosis result in diploid gametes instead of the normal haploid gametes

- Individuals with diploid gametes mate with each other and produce polyploid offspring

- Resulting individuals can not breed with progenitors (parents), but can breed with each other.
A single genetic event resulting in *instantaneous speciation* is an example of what type of speciation?
Polyploidy.

A type of hybridization speciation.
What are the 2 types of Allopatric speciation?
1) Vicariant Speciation

2) Peripatric Speciation