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115 Cards in this Set

  • Front
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Homologies

Similar structure or characteristics that are acquired from a common ancestor .

Homologous structure

Similar in fundamental structure,position and development. But....different functions e.g paradactyl limb

Divergent(adaptive) evolution or Divergence

Show evidence of evolution and similarities traced to a common ancestor. E.g Habitat and mode of life changed the structures

Analogous structure

Similarly functioned structure in different organism because they evolved in similar environments(not from common ancestors) e.g bat and Butterfly wings

Biological evolution

Genetic changes in a population (of a living organism) that are inherited over successive generations. Due to natural selection eventual lead to the formation of a new species

Macro evolution

Development of new life form or species from earlier life forms over many generations(extended geological period) I.e involves speciation


Usually can’t be seen happening in current time

Micro evolution

Change occluding not at a level of species. That’s not over a long period of time . I.e small changes in gene pool

Phylogeny

Scientific study of evolutionary relationships among species

Phylogenetics

The study of evolutionary relationships among organisms which have been discover through evolutionary lines of evidence

Phylogenetic tree

Show evolutionary relationships over time.


Usually Comprised of stasis, lineage splitting and adaptive radiation

Cladogebesis

Is an evolutionary splitting event where a parent species splits into 2 distinct species (forming a clade)


That live at the same time

Clade

Life-form groups consist of common ancestor and it’s descendants- representing single branch on tree of life

Anagenesis

Ancestral species gradually accumulate change over time. Until, species is sufficiently different enough from it’s origin.


Lineage in phylogenetic tree doesn’t split.


Ancestor does not exist at same time

Evolution vs natural selection

Evolution must involve a significant gain of genetic information. So that the common ancestor dies out. Whilst, natural or artificial selection always has a loss in vigour i.e only suitable for current conditions

Punctuated Equilibrium

Theory by Stephen J Gould and Niles Eldredge explain long period fossil record unchanged and absence of “missing links”


Species is In equilibrium untill rapid change “punctuated “ (interrupted) their existence

Gradualism

Darwin’s model of slow gradual change in a species

Natural selection

Seen as a “mechanism of evolution” were nature selects fittest individuals for survival


(best suited individuals adapt to and environment pass on favourable hereditary trait)


Occurs in populations not individuals

Artificial selection

Deliberately breed plant or animal to acquire desired characteristics- human is selective pressure


I.e new individuals not necessarily best suited for survival

Selective breeding(unnatural selection )

Specific organism with specific trait is bread with organisms of same characteristics- to eventually form desirable trait


E.g farmers/ breeders

Steps of Natural selection

1)Overpopulation,


2)Variation must be hereditary in individuals ,


3)Changes in environment,


4)Selection occurs(Best suitable individuals reproduce) others die out


If evolutionary ..


5)All individual in population have trait


6) Numerous changes (eventually lead to new species)



Selective pressure

Certain environmental factors exert pressure and trigger Natural selections e.g food competition , temperature change ect.

Examples of Artificial selection


Animals and agriculture

Canis lupus( grey wolf)- Canis Lupus familiaris( Domestic dogs)


Brassica oleracea(wild mustard)


- turnip, cabbage, broccoli... by out breeding

Inbreeding

Cross closely related individuals to maintains desired trait( increase homozygosity- identical alleles brought together more frequently )

Out breeding

Cross unrelated individuals( different strains, populations ect ) to introduce new or stronger traits. Offspring are genetically different


( Promotes heterozygosity-alleles are different )

Formation of a new species ( mechanism of evolution)

Natural selection, gene flow , genetic drift and Polyploidy( type of reproductive isolation )

Species

A group of organism that can interbreed to produce viable (fertile) offspring

Speciation

The evolutionary process whereby new species form-


E.g the formation of a species increases range of organisms on earth and biodiversity

Extinction

the permanent loss of a species ( results in decreases in biodiversity on earth )

Population

A group of organisms that live in a particular place at a particular time with the ability to freely interbreed.


I.e same species share gene pool but they can be free gene flow between populations

Gene pool

Total number of allele in all the reproductive individuals in a particular population

Gene flow

The exchange of alleles between populations


E.g migration and seed dispersal constitute


Im(in) or En(exit)-migration

Reproductive isolation

Isolation prevent ancestral and new species interbreeding


Can be geographical separated(allopatric) or occupy same area (sympatric)

Summary of speciation

1)Variation in a population


2)Evolution by Natural selection-“Adaptation to the niche”


3)Reproductive isolation prevents interbreed


4)New species formed

Types Variation

Continuous variation and discontinuous variation

Continuous variation

Range of Phenotypes for same characteristics.


Usually polygenic characteristics ( more allele = more possible combos)

Discontinuous Variation

Phenotypes fit into seperate characteristics- no intermediate form


Usually only single pair of allele occur(fewer possible combo)


E.g HBO blood groups, tongue roll ect

Mechanism of variation

Genetic Recombination


Mutations( gene or chromosomal)


-Variation is essential because environment selects best adapted from the variation

Mutation

Change in at-least one nucleotide base, usually due to the presence of mutagens


- only Gonad and Gamete mutation inherited

Fixed mutation

Once all organism of a population have new good allele

Chromosomal mutation

Chromosome structure or chromosome number change


Polysomy

Chromosome number more than normal diploid number

Polyploidy

Occurrence of multiple sets of chromosome- often in plants e.g tetraploid Duran wheat

Genetic drift

Random changes in the frequency of characteristics in a population. I.e role of chance


Important in very small population. More influential than Natural selection

Types of mutations( by effect)

Beneficial, Harmfully, Neutral

Contrast finches from Galapagos island to mainland finches

Island - Darwin saw approximately 13 different species [Each diff. Beak and food source ] + diff. To mainland


Mainland(1000s km away) - All seed eaters belonging to 1 species

How did variation occour in Galapagos finches?

1)Ancestral species( relatively few) arrive on floating trunk(drift wood) . Due to strong current.


2) little competition and lots of empty niches


3) mutations occurred as they multiplied( Variation in Pop.)


4) On Each island different trait considered beneficial mutation-Due to unique environment.


5)Finch with beneficial trait reproduce most successful. So, trait multiplies


6) mutations occouring constantly allows for the formation of new species - that fill a variety of niches


(E.g beak size differs for function)



Nb.Absence of other birds- (confirm Darwin’s theory- species change as environment change)


Finches Can’t interbreed cross islands or with mainland species



Pros and cons of In breading

Pro


-Predictable offspring


-undesirable trait can be isolated( often recessive gene)


Con


- offspring low resistance to disease


- high mortality rate and abnormality


-loss of Vigour(heterozygosity)


-gene pool gets smaller(possible extinction)


- recessive genetic disease prevalent

Human and animal example of inbreeding

Cheetah- Game reserve. Population genetically isolated. Increase misaligned jaws prevalence( less effective hunting)


- inbreeding can either reinforce or eliminate trait


Human- Presence of double recessive (e.g Tay-Sachs or haemophilia ) occurs


Due to small population, geographic isolation,royalty ect


Bottle neck effect

A Sudden decrease in population


causes a decrease in the gene pool


Causes a decrease in variation.


Often inbreeding happens afterwards-which further lowers variation


Causes: Humans(hunting), Natural disasters, disease ect

Pros and cons of Outbreeding

Pros


-Genetic variation, new trait introduced, bigger gene pool


- increase hybrid vigor


-reintroduce lost traits


Cons


- typical traits lost


- New races develop( loss of seperate racial breed)


-less predictable offspring

Animal examples of Outbreeding

Bonsmara cattle Breed


2 vulnerable cattle breed in subtropical RSA. So out-breeding had to occour.


Afrikaner ( irregular birth, poor meat quality, late sexual maturity)


Herelord & shorthorn(unable to adapt to climate, susceptible to tick related disease)


Bonsmara(Good meat and milk produced, regular birth, more disease resistant)


Bons(Prof. Jon Bonsma)


Mara(research farm used in Limpopo)

Founder effect

Loss of genetic variation when a new population is formed from a number of individuals moving to a new area


Nb=gene pool of new pop. contains genes of founder members

Examples of founders effect

-Afrikaans population(40 Dutch family 1652) Brought Huntington’s cholera


-Cheetah(likely due to catastrophic event)-

Fecundity

Ability to produce healthy offspring

Convergent evolution

Many unrelated organism independently develop adaptation for similar environment .


Think analogous structures

Divergent evolution

Single common ancestor gave rise to a variety of different, but related organisms


Think homologous structures

Mechanism(types) of speciation

Geographic(allopatric) and Sympatric

Allopatric speciation

New species originated from ancestral species -now separated by geographical barrier.


Often followed by , Reproductive Isolation.

Reproductive isolation

No gene flow occurs.


New mutation in each population lead to new allele forming and different gene pools.



Since environmental pressure different in each area. Natural selection occur independently.


Population Become Genetically different, Can’t interbreed( seperate species)

Example of allopatric speciation c

Galapagos finches(On different islands) In different niches .


Remember-


No need to fly since successful in differing niches.



Cambrian Explosion

Every phylum suddenly existed. (All complex life appeared)


Can’t be gradualism. That’s why they came up with punctuated equilibrium

Adaptive radiation

Burst of divergence from single lineage-giving rise to many species to fill niches


E.g Galapagos finches or protea in RSA AUS & SA


Usually allopatric speciation


Nb. Those who can experience adaptive radiation usually start with an unusual characteristic

Sympatric speciation

New species arise from ancestral species in same geographical area-


Thereby, Experience reproductive isolation suddenly.

Short examples of sympatric speciation

Polyploidy in plant( not evolutionary though)- can’t form matching homologous pairs


Hybrid species formation - In plants sometimes hybrid produces fertile offspring that is isolated( Not common in nature)

Cichlids(of lake Malawi) as an example of rapid sympatric speciation

Same lake with different selective pressures


1)Habitat specific selection- sandy(bottom) vs rocky( sides)


2)Food specific selection- few common ancestors ate great variety. Increase in pop. Lead to increase in interspecific comp. Develop specific food source niche(filled empty ones.)


3)Sexual specific selection- Original random mutate -bright color - proved beneficial( female liked them)



Nb. New species occur in same geographical area- No geographical barrier for reproductive isolation

Types of Reproductive Isolating mechanism( prevent gene flow)

Prezygotic ( occurs before fertilisation to prevent mating and/or fertilisation )


Postzygotic( occur after fertilisation and cause abnormal development of zygote-no viable offspring)

Prezygotic strategies of reproductive isolation

Temporal- Breed at diff. Times


E.g Nocturnal vs diurnal animal


E.g Rana Boylii (Feb) & Rana Aurora(April)


Behavioural- Courtship ritual species- specific I.e only species recognise(so save energy no mistake mate)


E.g Dogs and wolves courtship display differ


Mechanical- different plant in different adaptation (structure and colour scheme for pollinator)



E.g wind vs insect pollinated plants vs bird


Insect pollinated plant release pheromones to atract


Bird pollinates usually un scented and beak shaped


Reproductive- e.g mating calls(roar,hum,sing) or colour change indicating maturity

Post zygotic examples of reproductive isolation

Hybrid isolation


Development of sterile offspring e.g mule


show hybrid vigour(stronger than parents) - Faster than donkey and hardier than a horse

Classification of hominids

Class- Mammalia


Order- Primate


Super family-Hominodea


Family -Hominidae( other is hylobatidae)


Sub Family- Hominae (other is ponginae)


Tribe- Hominini( other is gorillini)


Species- Homo(other is pan)


Other grate apes


-Pongo- Orangatan


-Gorilla- Gorrila


-Pan-chimp



Extra - Hylobates-gibbon

Hominids

All members of family hominidae (great ape and human)


- some say not great apes only fossil ancestor

Hominins

Refer to Human and fossil ancestors ( hominini tribe without chimps)


- this context hominids are bipedal animal infossil record show mix of ape an human features

Anatomical Similarity between great African ape and humans

-Opposable thumb day is the the rèqqq pul( power grip but only human have done motor grip)


- 2 hands on each 5 finger + same with feet and toes


- long arms rotate freely at shoulder joint ( all direction movement)


-Flat nails


-No tail


- sexual dimorphism


- eyes have cones(and rods) so colour vision is possible

Anatomical differences between African ape and humans

Quadrupedal vs bipedal


Vertebral column: C shaped vs s shaped


Feet: Flat vs arched


Pelvis: Narrow long vs short wide


Forman magnum: at back of skull vs closer to front


Brain: Small undeveloped vs larger developed


Prognathism: High vs reduced


Jaw strength(zygomatic arch) : high vs reduced


Canines; prominent vs small,same size as other

Anthropology

The study of the origin and social relationships of human kind

Paleoanthropology

The study of human evolution by study fossils of humans and artifacts

Evidence of common ancestor for hominids

Fossil evidence(bipedalism, brain size, Crainial & Brow ridge, prognathism, dentition and palate shape )


Genetic evidence( mtDNA and Chromosomal DNA comparison of living Hominds)


Archaeological Evidence:tools(Oldowan, Acheulian, mousterian and recent)

Primate subdivision

Lemur, lorises,tarsier, New world primate(platyrrhini), humans , African ape and old wold primates(catarrhini)

Arboreal

Living in trees

Palaeontology

The study of plans and animal fossils

Archaeology

The study of artefacts left behind by early human populations

Hominid sequence

Ardipithecus ramidus (5.8-4mya)


Australopithecus (4.5-1.4mya) e.g laetoli footprint(3.3mya)


Paranthropus Robustus(aprox 4mya)


Australopithecus Africanus(3-2mya)


E.g Taung Child and Mrs Ples


Australopithecus afarensis (3.5mya)


Australopithecus prometheus


Australopithecus sediba(2-1.7mya)


Homo habillis(2.2-1.6mya)


Homo ergaster(2-1.4mya) e.g Tarkana boy


Homo erectus(1.8-0.3mya)


Homo heidelbergensis(700-200kya)


Home neanderthalensis(230-30kya)= not direct to man


Homo naledi(about 300kya) =not directly relate to man


Homo sapiens(200kya-now)


Sometime make us sapien sapien



Ardipithecus ramidus

Ape like everything -probably bipedal(still opposable toe)


Changed depending on climb or walk


1993- Afar valley

Australopithecus

Gracile and robust(Paranthropus robustus)


Form


First bipedal primates (non opposable toe and tenth humanish)- laetoli footprints show this


Not common ancestor as develop simultaneously to homo genius

Australopithecus africanus

1924-Taung child(3-4yr child skull) Raymond Dart, taung, NW of Kimberly


1947-Mrs Ples(Complete adult skull + Varius bones) Robert Broom, Sterkfonteon caves


Most plant-less meet

Australopithecus afarensis

1974-Lucy, Fossilised female skeleton(pelvis)


Face, limbs ape-like but show bipedalism


Donald johanson, Hadar afar region of Afar Valley

Australopithecus Prometheus

1994- Ankle foot bones by Ron Clark


1997- found rest still excavate now


Hand similar to modern human and bipedal


Sterkfontein caves

Australopithecus sediba

2008- lee Berger and Matthew find mother and boy(Karabo)


1st new hominid species since mrs Ples(announce 2010)


Most complete hominid fossil so far-


Consider transition fossils - A africanus and h sapien


Can Finally walk for long distance(pelvis like humans)


Prof Berger...Think Australopithecus could be related to homo

Difference homo and Australopithecus genus’s besides obvious ones

Homo brain- enlarged for language and use tools


Homo skull- more human like less prominent brow ridge and flat face

Homo habillis

Lived with Australopithecine(2.2-1.6mya)


Discovered in 1960( smaller body then Australopithecines)


Consider 1st use stone tools- larger brain so could use hands

Homo erectus

Closer to modern human than ape-man(1.8-0.3mya)


1st to use fire(larger Brain)


large body like humans


Face still slightly ape-like(brow ridge and jaw)


Out of Africa model- 1st exodus


Most complete- Tarkana boy(1.5mya), lake Tarkana-sometimes called homo ergaster


Pekeng man(0.4mya) China


Java man (0.7mya) Java, Indonesia

Homo neanderthelensis

230- 30kya


1856 Neander valley Germany discovered


Brain larger than Homo sapiens but limited development - buried dead


Prominent brow ridge and big nose

Homo sapiens

200kya-now


make tools from bone and stone( bow, fish hook(


Life in tents and formed community


Animal Migration followers and domesticate + cultivate land


Jewellery and body paint


Bury the dead

Homo Naledi

Found 2015 by cavers


Probably in-burial site- difficult getting in


Not directly related to humans extinct side branch


Found in dinaledi chamber(limestone caves)


About 1500 specimen and 15 individuals

Homo heidlbergensis

700-200kya


Often referred to as archaic homo sapien


1st species to group hunt and build simple wood+ rock dwelling


Taller than modern man , brain slightly smaller

The SAN the world oldest extant people

mtDNa show greatestbgenetic diversity


Suggested direct descendent if homo sapien before the migration


Taken of ancestral land by African and European population


I’m unfavourable condition now


100k left in Southern Africa (RSA,BW,Namibia ,Angola)

Interdependence on Hominid skills

Scientist differ explain how bipedalism, fire making, tool making , culture and brain capacity link


Difficult to distinguish which happened first. So, interdependent(Essential to each other )


Bipedalism- need tool skills to actually hunt event though see prey


Develop complex tool kit- hunt kill larger prey(more protein). Better for brain development


Increase brain capacity depend on improved diet- bigger brain =more like to get skills for fire making or tools to hunt


Develop fire making skills- greater range of food(Growth and development follow)


Hunting- require language and communication

Out of Africa hypothesis

Homo sapien originate in Africa 200kya and migrate relatively recently (50kya)


1st Exodus- Homo erectus left Africa 1.8mya establish population in Europe , Asia and Indonesia


2nd Exodus- 50kya Homo sapien also left Africa to out compete each life form they came up against.


( high level technical skill, better environmentally adapted )




Early Homo sapiens when to Africa and Asia to outcompete homo erectus

Multi regional Hypothesis

Homo Erectus left Africa & simultaneously across the world gave rise to Homo sapiens( parralel evolution)


Denosovan(Asia) Neanderthols(Europe)


Homo Erectus in both


Both model supported by genetic and fossil evidence

Evidence of African origin for all humans

Genetic links(DNA)


- Chromosomal- similar to great apes and great apes in Africa (rarely tested because degrades)


- Mitochondrial- simpler time analyse fewer genes code for about 13 proteins


Only received from mum( only nuclei of a sperm enter ova)


Compare different population for extent of variation( logic most variation=more time to mutate)


Mitochondrial eve- use expected rate of mutation /year, to find out lived 200-150mya (support out of Africa )


-Y chromosome- Only inherited by males


no crossing over and exchange material.


So, Pedigree form on male line , Lead to Y chromosome Adam


Was


fossil record (largest collection of hominid fossils in africa)


-Great Rift Valley(Kenya,Ethiopia, Tanzania)


Deep split in river due to shift in tectonic plate (valley divided in to western and eastern section)


Largest lake on content- probably attract lots of animal and hominids


Since 1993 Leaker family- contribute a lot work their


-Cradle of human kind(richest source hominid fossils globally)


1/3 of works holiday fossils


1998 - Prof Philip Tobias campaign- Unesco world heritage site


Consists of 13 fossil sites- Sterkfontein caves most known

Cradle of human kind

Back (Definition)

Great Rift Valley

Back (Definition)

Hominid fossils in SA

Taung, NC- A africanus( Raymond Dart and Philip Tobias)


Blombos,WC- h sapien , Art and complex hunting kit


Pinnacle Point, Mosael bay, WC- earliest use of fire and ochre artifacts


Klassie River caves, EC- h sapien burriel


Florid bad, FS- h heidelbergensis skull


Border cave, KZN- h sapien, tools and marked bones (Raymand Dart)


Makapansgat,LP- a afrikanus & h sapien(Raymond Dart)


Langaaban,WC- h sapien,” Eve’s footprints “

Dr Robert broom(1866-1951)

Mrs Ples( a africanus) in Sterkfontein cave 1947 & Australopithecus fossil around

Prof Raymond Dart(1893-1988)

Taung Child(small a. Africanus skull) in 1924

Proff Philip Tobias(1925-2012)

Valuable research In Sterkfontein Caves and identified homo habillis with Louis leaky in Olduvai Gorge

Proff Ronald Clark(1944-)

Discover little foot( a prometheus) in Sterkfontein cave -1994

Dr Andre Kruger (1938-2010

Discover complete scull female a robustus near Sterkfontein in 1994

Evolutionary trend in hominids

Quadrupedal- bipedal


Decrease


-Cranial and brow ridge size


-Size of teeth


-Prognathism


Increase


-Brain capacity


-Dexterity (skill in performing tasks especially with hands)

Swaartkrans

First site show 2 specie lives in same are at same time


A robustus and h erectus found


Bob Brian discover- evidence controlled use of fire(1mya)

Resistance ( in terms of evolution in present times)

Ability of an organism to reproduce in the presence of chemical which would normally limit growth or cause their death

Evolution in present time

Evolution usually Takes place over centuries and can be difficult to observe.


But, in organism that multiply rapidly( bacteria, virus and some insect and birds ) = possible)

Types of evolution in present tie

Resistant to insecticides in insects


- DDT is selective pressure( variation through random mutate)- used to control and exterminate mosquitoes


Resistant to antibiotics in bacteria



Drug resistant TB bacteria


- MDR TB( resist 2 AB) and XDR TB(resist all AB, must use chemotherapy but only cure 50%)


- That’s why DOTS( directly observed treatment short course) was invented

Evidence pronates close common ancestor

Arboreal- common ancestor probably lived in trees


- kept clavicle( stabilise shoulder and allow weight when hanging)


- opposable thumbs for grasping


enlarged complex brain relative to body size


- complex social behavior- usually extended care


- Eyes face forward and reduced snout- rely on stereoscopic vision not smell


Large percentage of shared DNA


- have full trichromatic vision

Adaption

Procces by which an organism becomes better suited to its environment

Clad a gram

Branching diagram showing classification relationship between spieces