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131 Cards in this Set

  • Front
  • Back
What is Ecology?
the study of the distribution and abundance of organisms and the factors and interactions that determine distribution and abundance
(where are they? how many are there? why?)
Where do the roots of modern ecology lie?
natural history, human demography, biometry, and applied problems of ag and medicine
-hunters & gatherers
-Aristotle
-Herodotus & Plato
What is Farr's Rule?
as the population density increases the death rate increases.
What are abiotic factors?
non-living chemcial and physical factors (temp, light, nutrients, water)
What are biotic factors?
living biological factors (other organisms, competition, predation)
What is an organism?
a single individual of a single species
What is a population?
individuals of the same species living in the same geological area
What is a community?
2 or more populations living in the same geological area
What is an ecosystem?
comprising the community together with with its physical enviornment
What is the mechanism for studying an ecosystem?
energy flow and cycling of nutrients among abiotic and biotic components
What is the mechanism for studying a community?
interactions among organisms
What is the mechanism for studying a population?
factors that affect population size and composition
What is the mechanism for studying an organism?
behaviors, environmental physiology, morphology
What is the mechanism for studying a tissue, cell, organelle, and/or molecule?
cell physiology, biochemistry
What are examples of ecological evidence?
-observation and monitoring in the natural environment
-manipulated field experiments
-controlled lab experiments
-math models
What is the goal of ecology?
to observe patterns, describe processes, and use this information to predict, manage, and control
What is a p-value?
(probability levels)
-measures the strength of conclusions being drawn
What is the null hypothesis?
assumes that there is no association between the two variables
Why does a species occupy a specific area?
-it evolved there
-it evolved somewhere else and dispersed to that area
Why does a species not occupy a specific area?
-it evolved elsewhere and never dispersed there
-it was once present but is no longer there
What are large scale distribution patterns (bio-geographic regions)?
-based on taxonomic similarities of organisms living there
-boundaries are set where species composition changes dramatically over short distances
What are large scale distribution patterns (climate)?
-average atmospheric conditions found over time (temperature, precipitation, wind velocity)
-varies because of differences in the amount of solar energy
What are large scale distribution patterns (biomes)?
-climate determines the distribution of biomes
-terrestrial biomes are based on the structure of their dominant vegetation
-aquatic biomes are based on their physical/chemical differences
-in each biome, vegetation has similar traits, but may not be evolutionarily closely related
What is a transplant experiment?
-the major conceptual techniques used to determine the factors limiting distribution
-move organism and see if it can survive and reproduce in the new environment
-follow through at least one generation
What does it mean if the transplant is successful?
the potential range of a species is larger than its actual range
What does it mean if the transplant is unsuccessful?
potential range of a species is the same as the actual range
If the transplant is successful, why might the species not live there?
-lack means of transport (dispersal)
-can move but choose not to (habitat selection- area is inaccessible or unrecognizable as living space)
If the transplant is unsuccessful, why can't the species survive there?
-limits imposed by other species (predators, competitors, parasites, disease)
-positive effects of interdependent species
-some other chemical/physical factor
What is dispersal?
the movement of organisms from one geographical location to another
What is the Southern California Intertidal Community example?
**dispersal**
hypothesis:
-barnacles can get south but die
-point conception is a barrier to larval dispersal
results
-barnacles survive in control and transplant.
*point conception is a barrier
What happened in the Tasmanian Blue Gum (Eucalyptus) example?
**dispersal**
-introduced in CA for railroad ties
-no native predators, out competes natives
-favorable abiotic enviornment
*barrier to dispersal: breached by human introduction
What happened in the Black rat and Mongoose example?
**dispersal**
-human introduction of black rats to hawaii (destroyed crops, native plants and native bird eggs)
-Mongoose introduced from India to eat rats (both nocturnal: eat birds and eggs)
*barrier to dispersal: breached by human introduction
What is habitat selection?
(factors limiting distribution)
-organisms can move but choose not to live in certain habitats
-distribution may be limited by the behavior of individuals in selecting their habitat
What happened in the Dragonfly example?
**habitat selection**
-males occupy territories they can defend
-determines where females will oviposit
-smaller than available reproductive habitats
*behaviorally contained habitats
What happened in the Tree Pipit and Meadow Pipit example?
**habitat selection**
-both are ground feeders and feed on the same organisms
-tree pipit is absent from treeless areas that the meadow pipit inhabits
-tree pipits breed only in areas having one or more tall trees
-meadow pipit has a similar song but it ends on the ground
*behaviorally constrained habitats
How are biotic interactions a factor limiting distribution?
-the local distributions of many organisms are limited by the presence of other organisms (competition, predation, parasites, disease)
What is competition?
(-,-) interaction
-can occur between 2 or more organisms or species that exploit the same types of limited resources and live in the same geological area
-intraspecific: competition within species
-interspecific: competition between 2 different species
What is interference competition?
direct physical interaction over resources
What is exploitative competition?
indirect interaction over resources
What are natural patterns indicating competition?
1. When species B is absent, species A lives in a wider range of habitats (competitive release)
2. If competition is strong, the geographic ranges of the two competitors may not overlap but have sharp boundaries.
What is predation?
(+,-) interaction
predation can limit the distribution of organisms:
-direct consumption of prey
-behavioral modifications of prey in the presence of predators
What are abiotic factors (physical/chemical) limiting distribution?
-temperature and availability of water
-organisms have an upper and lower lethal limits to physical and chemical factors
-solutions are either acclimination or adaption
What are abiotic factors (temperature) limiting distribution?
-cells freeze at 0 C and proteins denature at 45 C
What are abiotic factors (water) limiting distribution?
-water is essential for life
-availability varies greatly among habitats
-osmoregulation problems (lose or gain water)
What are abiotic factors (light) limiting distribution?
-provides the energy that fuels nearly all ecosystems (photosynthesis)
-the intensity and quality of light limit the distribution of photosynthetic organisms
-important to the development and behavior of many plants and animals that are sensitive to photo period
What are abiotic factors (soil) limiting distribution?
-the physical structure and nutrient content of soils (limits the distribution of plants and animals that feed on them)
-the composition of substrate (affect water chemistry of streams and rivers which influences the residence of plants and animals)
-in marine systems (soil structure in the intertidal zone and seafloors determine the types of organisms that can attach or burrow in those habitats
What are abiotic factors (periodic disturbance) limiting distribution?
-catastrophic disturbances (fires, hurricanes, tornadoes and volcanoes) can devastate organisms and limit distribution
-frequent disturbance: may be predictable and organisms may be evolutionary adapted to the disturbance
-infrequent disturbance: may be unpredictable and organisms will not have acquired the evolutionary adaption to them
What are abiotic factors (pH and salinity) limiting distribution?
-affects the distribution of marine and freshwater organisms
-distribution is limited by high and low pH
What are abiotic factors (currents and oxygen) limiting distribution?
-many organisms are dependent on wind and water currents to disperse their propagules or young
-oxygen can limit the distribution of organisms in lake
What happened in the large and small barnacle example?
-as adults, barnacles are found in discrete zones (but larvae are spread everywhere)
-move large high (upper limit= dessication)
-move small low (small lives if large is removed=competition)
What happened in the predatory snail and large barnacle example?
-snail and barnacle are found in discrete zones as adults (barnacle larvae settle in all regions)
-cage experiment:
in cage=barnacles lived
outside cage= barnacles consumed (predation)
What happened in the serpentine soil and goldfield flowers example?
-goldfield flowers are only found on serpentine soil and other plants are not found there
-other plants to serpentine=died (soil toxicity)
-goldfield to sandstone soil=died when there were competitor but lived when other plants were removed
What drives population dynamics (the change in population density through time and space)?
natality, morality, immigration and emigration
What is density?
the number of individuals per unit area or volume
What is density measured by?
-total counts (count all the individuals in the population)
-sub-sampling methods (estimate densities and total population size ex. mark and recapture)
-indirect indicators (number of nests, fecal droppings, tracks, ect)
What is the mark and recapture formula?
marked animals (2nd)=marked (1st)
total captured (2nd)=total population(N)
What is natality?
birth rate= number of offspring produced per unit time
-depends on organism
-birthrate= number of births/ size of reproductive population
What is morality?
death rate= number of offspring dying per unit time
-death rate= number of deaths/ size of population at risk
What is the immigration rate?
the number of individuals moving into an area per unit time
What is the emigration rate?
the number of individuals moving out of an area per unit time
Immigration and emigration are more important with....
smaller temporal/spacial scale
Birth/death rate are more important with...
larger temporal/ spacial scale
What drives population dynamics (the change in population density through time and space)?
natality, morality, immigration and emigration
What is density?
the number of individuals per unit area or volume
What is density measured by?
-total counts (count all the individuals in the population)
-sub-sampling methods (estimate densities and total population size ex. mark and recapture)
-indirect indicators (number of nests, fecal droppings, tracks, ect)
What is the mark and recapture formula?
marked animals (2nd)=marked (1st)
total captured (2nd)=total population(N)
What is natality?
birth rate= number of offspring produced per unit time
-depends on organism
-birthrate= number of births/ size of reproductive population
What is morality?
death rate= number of offspring dying per unit time
-death rate= number of deaths/ size of population at risk
What is the immigration rate?
the number of individuals moving into an area per unit time
What is the emigration rate?
the number of individuals moving out of an area per unit time
Immigration and emigration are more important with....
smaller temporal/spacial scale
Birth/death rate are more important with...
larger temporal/ spacial scale
What methods are used to identify how population size changes (dynamics)?
-direct observations (not always practical)
-view age structures (life tables and survivorship curves)
-math models (
What is the formula for the simple population growth model?
Nt=No+B-D+I-E
What is needed for exponential growth?
unlimited resources!
dN/dt =rN

(r=b-d)
What happened in the Northern Elephant seal example?
**exponential growth model**

-close to extinction
-hunting stopped: habitat and resources were abundant
-exponential growth
What happened in the European Rabbits example?
**exponential growth model**
-rabbits from England introduced to South Australia
-each female can produce 6 liters of 10 rabbits each year
-population increased exponentially
-largest fence ever built failed
What is the carrying capacity(K)?
the maximum population size that an environment can support
-varies over space and time
What is the logistic growth model?
dN/dt=rN((K-N)/K)
How well does the logistic model fit the growth of natural populations?
-populations of very small organisms fit fairly well
-many populations do not stabilize at K and deviate from curve
-predictions are only correct when:
environment is constant, no predators, and no competition from other species
Why does the logistic curve not fit well for most natural population growth patterns?
because it assumes that:
-each individual added to population has the same negative effect on population growth (ex. flamingos and isolated plants)
-population approaches K smoothly (time lag)
-populations are large and density is important in regulation (ex. insects and microorganisms)
What is density dependent control?
factors that alter per capita birth/death rate in a population are dependent on population density (can stabilize population density)
-ex. parasitism, predation, competition can be density dependent
What was the bubonic plague example?
**density dependent control**
-blood pathogen that exists in rats.. transmitted by fleas
What is density independent control?
factors that alter per capita birth/death rates in a population are independent of population density (does not stabilize density)
-ex. physical/chemical factors: weather, drought, freezes, flood and fire
What happened in the monarch butterfly example?
**density independent control**
-migrate from canada to mexico
-logging removed temperature buffer
-freeze killed millions of monarchs
What is a stable population?
(birth rate=death rate)
-if perturbed population will return to initial density
-stabilizing forces dampen population fluctuations
-density dependent control
What is an unstable population?
-if perturbed population may not return to initial density
-destabilizing forces enhance population fluctuations
-inverse density dependents (long time lags)
What are metapopulations?
population is divided into discrete sub-populations which are connected by immigration and emigration
-growth and reproduction within patches
-migration between patches or colonization of empty patches
What happens to metapopulation when migration increases, decreases and remains constant?
-increases: local population fluctuations are dampened
-decreases: local population fluctuations are enhanced (increases probability of extinction)
-intermediate: shifting mosaic of occupied and unoccupied patches (persistence)
What is the metapopulation dynamics model?
(how does the fraction of occupied sites (f) change with time?)
f=fraction of sites occupied (proportion of patches that contain populations)
if f=1 all sites are occupied
if f=0 all sites are unoccupied (extinct)

df/dt=(probability of local colonization)(1-f) - (probability of local extinction)(f)

I>E then f increases
What happened in the Bay checker spot butterfly example?
**metapopulations**
-caterpillar larvae feed on specialized plants that grow on serpentine soil
-in drought years, most of these plants die, kills larvae
-one drought, 3 metapopulations became extinct
-largest subpopulation became new colonists
What is mutualism?
(+,+) both species benefit from the association
-ex. large African mammals & tick birds and sea anemone & zooxanthellae
What is commensalism?
(+,0) one species benefits from the association, the other is unaffected
-ex. shark & remora and cockroach & human
What is competition?
(-,-) both species are harmed from the association
-ex. lion and hyena
What is predation?
(+,-) one species benefits from the association and the other is harmed
-ex. humans & parasites (parasitism), tapeworms, and intestinal round worms
What is the Lotka-Volterra competition model?
dN1/dt1=r1*N1((K1-N-alpha*N2)/K1)
this gives the population growth of species one in competition with species 2
What are possible outcomes of competition?
1. both species coexist (stable or unstable equilibrium)
2. species one becomes extinct
3. species two becomes extinct
How do theoretical models for competition apply to natural systems?
ex. Gause
-competition between 2 species of yeast
-grew populations in isolation (logistic growth)
-then grew 2 populations together
-species 1 growth unchanged (logistic) and species 2 went extinct
*fit data to lotka-volterra equations
-species one had a higher competitive effect (beta>1)
-species 1 produced more alcohol and waste then species 2
What is the competition exclusion principle?
no two species can occupy the same ecological niche simultaneously (complete competitors can not coexist)
What is a niche?
the biological role of an organism within an environment
What is a fundamental niche?
the multitude of conditions in which an organism can survive and multiply
What is a realized niche?
where an organism actually exists due to ecological constraints
What happened in the parasitic wasps example?
**competition- competitive exclusive**
-competitive exclusion displaced prior species of wasps but now 2 species coexists but in different habitats in So Cal
Why doesn't competitive exclusion occur?
1. unstable environments (never reach equilibrium, low density of organisms)
2. environments in which species do not compete
3. fluctuating enviornments that reverse the direction of competition before extinction occurs
What are the four types of predation?
-herbivory
-cannibalism (KESHA)
-parasitism
-carnivory
What is the formula for the Lotka-Volterra Predator-Prey model?
prey population growth:
dN/dt=rN-(aPN)
predator population growth:
dP/dt=faPN-qP
What are the predicted outcomes of the Lotka-Volterra Predator-Prey model?
-equilibrium
-oscillations
-extinction
What happened in Gause's Laboratory Predator Prey example?
D(predator) and P(prey)
-D exterminated P and D died of starvation
-no oscillations
What happened in Utida's laboratory predator prey example?
azuki bean weevils (host) and parasitic wasps (parasite)
-interaction produced cycles (112 generations and 14 complete oscillations)
-oscillations converged and wasps died out
What happened in the Sea Lamprey example?
-canal linked Great Lakes to Niagara Falls
-first lamprey discovered in Great Lakes
-decimated lake trout
-lamprey control efforts (dams)
-trout population increased
What happened in the snow shoe hare and lynx example?
-hare decline due to decrease in birth/growth rate and increase in morality (predation)
-increased number of hares decreases food supply, increases exposure and predation
-as predation increases, increases growth/birth rate and decreases hare population
What are community characteristics?
-trophic structure
-species diversity
-dominance
-growth form and structure
What is the individualistic hypothesis (community)?
communities are chance assemblages of species with similar abiotic requirements
What is the interactive hypothesis (community)?
communities are assemblages of dependent closely linked species
What is species diversity?
richness(number of species in a community) + evenness(relative abundance of species in a community)
What is the Shannon Weaver (Wiener) Diversity Index?
H= -sum(Pi*ln(Pi))

Pi= proportion of individuals contributed to total abundance
The wider the average niche breadth, the...
fewer the number of species in the community (extinction via competitive exclusion)
The narrower the average niche breadth, the....
more species in the community (coexistence via resource partitioning)
How does intraspecific competition affect the short and long run?
short- decreases diversity (ecological)

long- increases diversity (evolutionary)
What is a keystone species (competitor)?
a species influences on the community is greater than expected that would be expected on the basis of its abundance
What happened in the keystone Starfish example?
-removed starfish from patches for 5 years
-space is a limiting resource (28 species coexist)
-in year 3: only barnacles and mussels left
-in year 5: only mussels present
-mussels are competitive dominant (excludes others from space)
*predator mediated coexistence
What is a disturbance?
an event that removes organisms that alters resource availability
-small scale (tree falls, rock flipped over)
-large scale (volcano, fire)
What is the intermediate disturbance theory?
1. When a disturbance is severe and frequent community is exposed of good colonizers with high reproductive rates
2. When a disturbance is mild and rare community is composed of good competitors
Species diversity is highest when disturbance is....
intermediate
-allows both colonizers and competitors to co exist
What happened in the Ellwood Boulder Field example?
-boulders of intermediate size had more species than small or large boulders
-glued small boulders in place (reduced disturbance)
-glued small boulders had higher species richness than loose small boulders
What is succession and what are the two types?
-transition in species composition over time
-primary succession=(bare soil, no organisms)
-secondary succession=(soil, some organisms)
What is the succession pathway?
1. early successional communities (r selection)
-good dispersers and high reproductive rates
2. late successional communities (K selection)
-good competitors and exist near carrying capacity
What is the equilibrium theory of island biogeography?
species richness is positively correlated with island size and negatively correlated with distance from mainland
What happened in the island biogeography example?
Flordia- 6 islands killed anthropods

-equilibrium within a year
-furthest island from mainland had fewest species
-equilibrium numbers similar between pre and post fumigation but species composition was different
What is the equilibrium view?
1. communities are structured by biotic interactions (competition, ect)
2. communites display global stability
3. niche diversification determine species diversity
What is the non-equilibrium view?
1. communities composition is always changing and is never in balance
2. no global stability
3. communities are patch works of disturbance. colonization and dispersal determine species diversity