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90 Cards in this Set

  • Front
  • Back
What are the essential rates?
growth, birth, and death rates
List the common ways to sample a population.
Quadrat survey
Random walk
Line transect
Point survey
Mark and recapture
Traps and passive collecting
When are quadrat survey's used?
Spatial subsampling, most often used with plants or fixed things such as corals, nests, burrows etc.
What is a nested design?
When a smaller quadrat is placed within a larger quadrat.
What are the most common sized quadrats and when are they used?
0.1 ha used for quick
and dirty forest census

5m quadrat used for rocky intertidal surveys, and seedlings.

1m quadrat used for intertidal and herbaceous plants including seagrass
How do you know what sized quadrat to use?
When the species diversity levels off
What type of sampling is a "random walk/swim"?
Spatial subsampling.

Predetermined distance and compass bearings (random).

Document target organisms encountered.
What makes up a point survey?
Spatial subsampling

Randomly selected points (or could be regularly spaced along line transect).

Document all target organisms within given radius (varies according to organism).
What is the formula for population index?
P= (100 NA)/L,
where:
P = population index,
L = length of transect in meters
A = size of flight area in hectares, and
N = butterfly numbers per 100 m of transect.
How is a point-quarter survey used?
Orientation of one axis can be chosen by random number (0-180o). Then distance to nearest individual is measured in each quarter.
What is the "Mark and Recapture" method?
Catch N number of a population at time 1.
Mark them (paint, clipped toes, tags)
Return at time 2 and resample same number from population and count the number (M) that are marked (i.e. recaptured)
What is the formula for total population size?
Total population P = na/r
where
P = total population
a = marked
n = total recaptured
r = marked recaptured
What are the assumptions for the Mark and Recapture method?
marking doesn't affect animals (behaviorally, physiologically, or ecologically);
marked animals are completely mixed in population;
probability of capturing a marked animal is the same as capturing an unmarked animal (closed population);
marked animals don't lose their marks;
marked mix naturally with unmarked.
List passive types of collecting population samples.
1. Pitfall traps for small reptiles and beetles
2. Cover boards
3. Light/black light trap
4. Plankton sampling
5. Soil fauna sampling using Berlese funnels
List the more common types of experiments.
Exclosures
Reciprocals
Additions
Growth responses
How do you test exclosure experiments?
Trying to measure the effect that one organism has on a system. Try getting rid of the organism.
Hunt them out and then keep them out using Exclosures.
Then monitor changes in other populations or habitat.
How do you test reciprocal experiments?
Observation: The same species of plant grows in two locations, but the ones at location A do better than at B.
Q: Is this pattern because the popn at A is better suited to conditions (evolutionary adaptation) or because location A is a better site?
Can test this by taking members of popn A and planting them at B, and taking some from B and planting at A. What happens?
How do you sample at large scale?
Data mining and using GIS (Geographic Information Systems )
What is the sequence of study?
1. Research the topic: what has already been done and what questions raised?
2. Formulate research question.
3. Decide if you are going to look for correlation or causation.
4. Statistics that you apply can then be hypothesis testing (ANOVA, MANOVA) or descriptive/hypothesis generating, e.g. Multivariate analysis and modeling.
Why are descriptive statistics needed?
They identify pattern in complex data that allow better hypothesis formulation.
In multivariate analysis, what is ordination?
Ordination: Treating samples as lying along continuous gradients. More realistic but a bit harder to understand at first.

-Performs similarity analyses and then maps the data points so that similar samples are clustered together, but segregated from those that are dissimilar.
In multivariate analysis, what is classification?
Classification: Dividing things into groups. Very intuitive as that is how our brain works, and can be useful to help us think about a problem, but artificial.
What is a community by definition from Clements theory?
It acts as a super organism
What is a community based on Gleason's theory?
It acts from individuals responses, not as a whole group.
What are the types of ordination?
Principal components analysis (PCA)
Detrended correspondence analysis (DCA)
Canonical correspondence analysis (CCA)
Non-metric multidimensional scaling (NMDS)
How are the various pools of Nitrogen cycled on the earth?
Pools of Nitrogen:
- atmosphere is 70% N, but not available for uptake
- N in sediments and rocks , but not available
- inorganic N in ocean is another large pool
- organic pools in plants and soils actively cycled
Describe the types of biological fluxes of nitrogen
Fluxes: N moves between biosphere and atmosphere
- biological: fixation, denitrification, nitrification
Describe the types of abiotic fluxes of nitrogen.
- abiotic: industrial fixation, lightning fixation,
fossil fuel and biomass burning, deposition
What organisms fix nitrogen?
-Heterotrophic bacteria that get organic C from environment and where N is limiting (e.g., decaying logs)
-Rates low due to low C supply and lack of O2 protection (0.1-0.5 g-N m-2 y-1)
-Also, cyanobacteria (free-living photo-autotrophs); symbiotic lichens (cyanobacteria with fungi offering physical protection)
***NO PLANT FIXES NITROGEN we talk about N-fixing plants, but this is always a symbiont associated with the plant, not the plant itself.
Why aren't plants labeled as N fixers?
Because microbial symbiont resides in root nodules, the plants don't do it themselves.
What is primary succession?
Succession refers to more or less predictable and orderly changes in the composition or structure of an ecological community. Succession may be initiated either by formation of new, unoccupied habitat (e.g., a lava flow or a severe landslide) or by some form of disturbance (e.g. fire, severe windthrow, logging) of an existing community. Succession that begins in areas where no soil is initially present is called primary succession, whereas succession that begins in areas where soil is already present is called secondary succession.
Is nitrogen an open or closed cycle?
N cycles as a closed system….much more cycles internally than flows in. Oceans are the major player.
Who fixes nitrogen more, nature or people?
PEOPLE! OH NO!
What happens if you "open" a closed system?
‘Opening’ a closed system means that the nutrient is now lost from the system..often washing out in streamwater or being volatalized as smoke Particularly true of N in fire-prone systems).
What are "Redfield ratios?"
C:N:P
106 C : 16 N : 1 P
What nutrient is typically the limiting nutrient in most terrestrial systems?
Phosphorus!

Most lakes are p-limited too
What happens if phosphorus increases in a water system?
Redfield ratios with alter and we will see algal blooms.
As the system degrades further it may become a bacterially-dominated rather than algal community…At that point you have a dead zone.
What is the ratio of carbon in soil, vegetation, and atmosphere?
Soils hold ~3 times as much carbon as vegetation or about twice as much as in the atmosphere.
What is soil made of?
Mineral matter (e.g. silica, aluminum, iron)
Organic material (decomposing squirrels etc)
Air
Water
Biota
What 5 factors does soil formation depend on?
1) Topography
2) Parent material
3) Climate
4) Biotic processes
5) Time
What 4 components are released in soil from weathering?
1) Minerals in solution (cations and anions), the basis of plant nutrition.
2) Oxides of iron and alumina (sesquioxides Al2O3, Fe2O3).
3) Various forms of silica (silicon-oxide compounds).
4) Stable wastes as very fine silt (mostly fine quartz) and coarser quartz (sand).
How do air spaces benefit soil composition?
Air spaces in soil are essential for keeping roots well supplied with oxygen.
How do air spaces benefit degrade soil composition?
Air spaces can fill with water. If filled for too long the soil becomes anoxic or waterlogged
How does soil pore size contribute to water distribution?
Large pores drain quickly (gravitational water).
Smaller pores provide water to plants.
Where do soils derive their mineral nutrients from?
Decay of parent material
Groundwater
N-fixing.

Cations are charged particles that are held on the surface of clays.
What is the cation exchange capacity (CEC?)
The degree to which a soil can adsorb and exchange cations.
In general does CEC decrease or increase with increasing soil acidity?
Decrease!!
How does CEC work?
When the cations are present in equivalent amounts, the order of strength of adsorption is:
Al3+ > Ca2+ > Mg2+ > K+ = NH4+ > Na+

So under normal conditions K + is more readily displaced (ie goes into solution) than Al3+

Plants use this to free cations from clays for absorption by pumping H + ions out of their root hairs.

If the soil becomes flooded by H + ions then this distorts the exchange and even H + ions can then displace Al3+.
What role does soil animals play?
Decomposition (shredding residues)

Mixing soil (aeration)
Does organic matter decay faster in boreal, temperate, or tropical forests fastest?
Tropical!
What 2 laws of thermodynamics govern ecosystems?
1. Energy can neither be created nor destroyed, only transformed from one type to another.
-conservation of energy
2. In any transfer of energy, some energy is lost.
-entropy
What are autotrophs?
Autotrophs: convert CO2 to organic carbon, i.e. all green plants and algae.
Variants being photoautotrophs (photosynthetic) and chemoautotrophs (deep sea vents, some bacteria).
What are heterotrophs?
Heterotrophs: Obtain carbon in organic form (predators or detritivores), i.e. all animals, fungi
What defines primary production and its producers?
Primary production – capturing light energy and storing it in chemical bonds of carbon compounds
primary productivity – the rate of primary production
primary producers – photosynthetic autotrophs
How do plants use the product of photosynthesis?
1. new proteins, tissues, cells, structures – growth & reproduction
2. fuel for the above processes
What is gross primary production?
Gross primary production (GPP) – total amount of energy assimilated by photosynthesis
What is net primary production?
Net primary production (NPP) – energy actually stored as biomass
How do you measure NPP?
energy per unit area per year
kJ per m2 per yr, or W per m2

1 g C assimilated = 39 kJ energy
can use plant biomass or CO2 uptake as an estimate of energy

Ignoring roots – annual aboveground net productivity (AANP)


Can measure via standing biomass or by CO2 usage.


Measure amount of CO2 absorbed by leaves
extrapolate from a very small area

Use radioactive isotopes to measure total uptake of C

In aquatic systems, measure changes in O2 concentration
What can limit primary production?
Light, temperature, water/nutrients
What is the energy transfer rate in a food chain?
With each step in the food chain, 80-98% of energy is lost
What is Ecological efficiency?
proportion of the biomass of one trophic levels transformed into biomass at the next higher trophic level.

Ecological Efficiency = Exploitation efficiency x Assimilation efficiency x Net production efficiency
What is Exploitation efficiency?
proportion of production on one trophic level that is consumed by the next higher level
What is Assimilation efficiency?
proportion of ingested energy actually absorbed by the body
EX:
seeds – 80%
young vegetation – 60-70%
grazing/browsing – 30-40%
wood – 15%
animals – 60-90%
What is Net production efficiency?
Net production efficiency = (biomass production)/(assimilated energy)
the proportion of energy not used for maintenance and not lost as heat
What is Gross production efficiency?
Gross production efficiency = (biomass production)/(ingested energy)
Why is Assimilation efficiency of herbivores is only 30-70%?
most plant tissue is not digested (cellulose, lignin, defensive compounds) by animals and ends up as detritus
What are the 2 independent food chains?
Herbivores and detritivores
What is Ray Lindeman known for?
Theorizing that 10% of energy flows from one trophic level to the next (Gross energetic efficiency). Untrue!
What is Residence time?
Residence time – average time that energy spends on one trophic level
= (energy stored in biomass)/(net productivity)
What is the Biomass accumulation ratio?
residence time based on biomass rather than energy
= (biomass)/(rate of biomass production)
What is the energy passing rate for homeotherms? Poikliotherms?
Homeotherms: 2-5%
Poikliotherms: 20-30%
What are the 3 types of food webs?
Connectance, energy, and redundancy.
What is a Connectance food web?
proportion of the potential links in the foodweb that are actually realized.
What is an energy food web?
web shows the relative flow of energy in the web
What is a Redundancy food web?
the number of links in the food web that perform the same ‘function’; complex foodwebs should have greater redundancy.
What is the prey to predator ratio in a balanced ecosystem?
Empirical studies show that the proportions of prey to predators is generally within the range of 2:1 to 4:1.
What is omnivory in a food web?
Omnivory is defined when a species is feeding on more than one trophic level.
What are the 5 indirect interactions in a food web?
1. Interference
2. Exploitation
3. Interspecific
4. Trophic cascade
5. Apparent competition
What is the Optimal Foraging Theory (OFT)?
-Predators will evolve to hunt in the manner most efficient to their physiology.
-Energetic argument: costs of hunting must be less than energy gain from feeding.
Costs are:
1. Hunting (from onset to capture)
2. Handling (from capture to digestion)
What is Dominance?
Dominance = basal area or aerial coverage, species A / area sampled
What is Relative dominance?
Relative dominance = basal area or coverage, species A / total basal area or coverage of all species.
What is frequency?
Frequency = intervals or points where species A occurs / total number of sample plots or points.
What is relative frequency?
Relative frequency = frequency value species A/ total frequency all species.
What is the importance value?
Importance value=
relative dominance + relative frequency + relative density*
What is Simpson's dominance index?
Measures the probability that two individuals randomly selected from a sample will belong to the same species


This index goes from 0 to the total number of species. An index of 1 is when all individuals belong to 1 species. When D ≈ 0 every individual belongs to a different species

D = SUM(Pi^2)
D = SUM ni(ni-1)/N(N-1)


Where D = Simpson’s dominance, between 1 and 0
ni = number of individuals in a given species (species i)
N = total number of individuals in the sample (community)
pi = proportion of individuals of a given species (species i)
S = number of species in the sample (community)
What is alpha diversity?
(within an ecosystem)
Species richness: number of species in a community
Species evenness: the relative abundance of individuals among species
Species dominance: measure of predominance of one or a few species in a community
What is beta diversity?
(between/among ecosystems)
Dimensionless number expressing the number of species gained going from one habitat to the next.
What is gamma diversity?
(within a region)
Dimensionless number expressing the diversity of habitats within a region, or a measure similar to b -diversity but across large geographic distances
Also defined as Geographic-scale species diversity
What is the Shannon-Weiner Diversity Index?
Similar to Simpson’s index except assumes an infinite number of individuals in the habitat.


H = Shannon-Weiner index
pi = proportion of individuals of a given species (species i)
S = number of species in the sample (community)
ni = number of individuals in a given species (species i)
N = total number of individuals in the sample (community)
What is Evenness?
Evenness is a measure of how similar the abundances of different species are.

Hmax = lnS ; E = H/Hmax

1>E>0