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35 Cards in this Set

  • Front
  • Back
Population Ecology
- Species distribution is partly determined by how fast the individuals can reproduce, how many of them survive and whether they disperse far away or not
- These are population processes, and they are determined by:
> fecundity (able to produce offspring)
> survival/mortality
> growth rates
> patterns of spatial distribution
53.1 Biological Processes influence pop. density, dispersion, and demographics

- Population
- Population is a group of individuals of a single species living in same general area
- rely on same resources, likely to interact and breed w/ each other

- Pop. described by their boundaries and size (number of individuals living w/in an area)
- may be natural boundaries (islands, mountains)
53.1 Biological Processes influence pop. density, dispersion, and demographics

- Density and Dispersion
- Density: number of individuals per unit area or volume
(oak trees per km in MN)
- Dispersion: pattern of spacing among individuals w/ in the boundaries of pop.
53.1 Biological Processes influence pop. density, dispersion, and demographics

- Density: a dynamic perspective
(counting pop. size, sample techniqe, addition, removal)
- pop size and density can be determined manually by counting.
- large mammals can be counted by plane.
(both impractical or impossible)
- Sampling techniques estimate density
> count number of oak trees in 100x100m plot, and avg plot, extend to area.
> accurate when sample plots are fairly homogenous

- Ecologists use mark-recapture method to estimate size of wildlife population

- Density changes as individuals are added or removed from pop.
- Addition occurs through: births or immigration
- Removal occurs through: deaths or emigration
53.1 Biological Processes influence pop. density, dispersion, and demographics

- Patterns of Dispersion
> Clumped, groups, uniform, random)
- Diff in local density important for ecologist to study, provide insight to associations, social interactions

> Clumped: individuals are aggregated in patches
- most common
- plants and fungi clumped on same log, insects and salamanders under same log due to humidity

> Groups: also increase effectiveness of predation or defense (hunting in wolf pack)

> Uniform: evenly spaced dispersion may be from interaction between individuals in pop.
- animals exhibiting territoriality (more rare than clumping)

> Random distribution: unpredicted spacing
- individual in pop is independent of other individuals
- strong attraction or repulsion from others
53.1 Biological Processes influence pop. density, dispersion, and demographics

Demographics
- Demography
Factors that influence pop density and dispersion patterns:
- ecological needs of a species
- structure of the environment
- interactions among individuals w/in a pop.

- Demography: Study of vital statistics of pop and how they change over time.
(birth rates, death rates)
- Way to summarize some vital statics of a pop, make a life table
53.1 Biological Processes influence pop. density, dispersion, and demographics

Demographics
- Life Tables
- Age-specific summaries of survival patterns of a pop

To construct a life table:
- follow fate of the cohort, group of individuals of same age, from birth until all individuals die
- Determine number of individuals that tdie in each age-group
- calculate proportion of cohort surviving from one age class to the next.
53.1 Biological Processes influence pop. density, dispersion, and demographics

Demographics
- Survivorship Curves
- graphic method to represent data in a life table> Survivorship curve
- proportion or numbers in a cohort still alive at each age.

Can be classified in 3 general types
1) Type 1: (humans) flat to start, then drops off significantly near the end> indicates death rates are low at young age, then increases rapidly as life goes on

2) Type 2: (rodents) intermediate, constant death rate over lift span. Diagonal line down

3) Type 3: (fish) curve drops sharply at start, high death rate. flattens out as death rates decline.
53.1 Biological Processes influence pop. density, dispersion, and demographics

Demographics
- Reproductive Rates
- Demographers who study sexually reproducing species, study the female, not male
- way to describe reproduction patterns of a pop is to ask how reproductive rates varies w/ ages of females.

- Reproductive table or fertility schedule is an age specific summary of the reproductive rates in a pop
- Constructed by measuring reproductive output of a cohort from birth until death.
53.2 Exponential model describes pop grown in an idealized unlimited environment

Per Capita Rate of Increase
- ideal condition: unlimited environment, no restriction: addition w/ birth and immigration, reduction w/ death & emigration
- Change in pop size during a fixed time interval w/ verbal equation:

Change in pop size = births + immigrants - deaths - emigration
53.1 Biological Processes influence pop. density, dispersion, and demographics

Per Capita Rate of Increase
- Math
N = population size
t = time
change N = change in pop size
change t = is time interval (appropriate to life span)

B = births in the pop during time interval
D = number of deaths

change N/change t = B - D
53.1 Biological Processes influence pop. density, dispersion, and demographics

Per Capita Rate of Increase
- per capita birth rate
number of offspring produced per unit time by an avg member of the pop.
- 34 births per year in pop of 1000
- annual per capita birth rate is 34/1000 or 0.034

annual per capita birth rate (symbolized by b), can use formula B=bN to calculate number of births per year

B=bN
B=0.034 x 500
B = 17 per year
53.1 Biological Processes influence pop. density, dispersion, and demographics

Per Capita Rate of increase
- Per capita death rate
m = mortality, allows us to calc expected number of deaths per unit time in a pop.

D = mN

If m=0.016 per year, we would expect 16 deaths per year in a pop of 1000 individuals

change N / change t = bN - mN
53.1 Biological Processes influence pop. density, dispersion, and demographics

Per Capita rate of increase
(diff between per capita birth and death rate
difference is per capita rate of increase, or "r"

r = b - m

- r indicates whether a given pop is growing (r>0) or declining (r < 0)
- Zero Pop. Growth (ZPG) occurs when the per capita birth and death rates are equal (r = 0)
53.1 Biological Processes influence pop. density, dispersion, and demographics

Per Capita Rate of Increase
- new equation for change in pop size
Can now rewrite equation for change in pop size as

change N / change t = rN

this equation is for a discrete, fixed, time interval (one year), doesn't include immigration and emigration.
53.1 Biological Processes influence pop. density, dispersion, and demographics

Per Capita Rate of Increase
- instantaneous growth
instantaneously, growth rate at a particular instant in time

dN / dt = r (inst)N

- r(inst) = instantaneous per capita rate of increase.
- time intervals change t are very short and are expressed in equation as dt.
- change t becomes shorter, discrete r approaches the instantaneous r(inst) in value.
53.1 Biological Processes influence pop. density, dispersion, and demographics

Exponential Growth
Pop increase under ideal conditions is called: exponential pop growth
- per capita rate of increase may assume max rate for the species, denoted as r(max)

dN / dt = r(max)N

Size of a pop that is growing exponentially increases at a constant rate, resulting in a J-curve.
53.3 Biological Processes influence pop. density, dispersion, and demographics

Logistic model describes how a pop grows slowly as it nears its carrying capacity
- Carrying capacity: symbolized by k, as the max pop size that a particular environment can sustain.

- carrying capacity varies over space and time w/ abundance of limiting resources

- if individuals can't obtain sufficient resources to reproduce, per capita birth rate (b) will decline
- if they can't consume enough E, or if disease hits (m) may increase.
- decrease in (b) or increase in (m) results in a lower per capita rate of increase (r).
53.3 Biological Processes influence pop. density, dispersion, and demographics

Logistic Growth Model
- Logistic population growth model, per capita rate of increase approaches zero as carrying capacity is reached.

- take pop. growth model, add an expression that reduces per capita rate of increase as N increases

- If carrying capacity is K, then K - N is number of additional individuals environment can support,

- (K-N) / K is fraction of K that is available for pop growth.

- multiplying exponential rate of increase r(max)N by (K-N) / K, we modify change in pop size as N increases:

dN / dt - r(max)N * (K - N) / k

- When N is small compared to K, term (K-N) / K is close to 1, per capita rate of increase r(max)(K-N) / K approaches max rate of increase
- When N is large and resources are limiting, (K - N) / K is close to 0, per capita rate is small.
- N = K, pop stops growing
53.3 Biological Processes influence pop. density, dispersion, and demographics
- Growth of lab pop. like beetles, and crustaceans fit an S curve well grow in an environment lacking predators.
- The logistic model assumes that pop. adjust instantaneously to growth and approach carrying capacity smoothly.
- often a delay before neg effects of an increasing pop are realized.
- if pop drops below carrying capacity, there will be a delay in pop. growth until increased number of offspring are actually born.
- logistic model is based on an assumption that regardless of pop density, each individual added to pop has same neg effect on pop growth rate.
- Allee effect: individuals may have more diff time surviving or reproducing in small pop (eg, single plant may blow away if alone, in a clump, plant is protected)

- Model is important for conservation in bio, for predicting how rapid a pop might increase after it has
- Conservation biologists can use the model to estimate the critical size shown which pop of certain organisms, may become extinct.
53.4 Life History Traits are products of Natural Selection
- Natural selection favors traits that improve an organisms chances of survival and reproductive success
- Trade off between survival and reproductive traits (freq of reproduction and number of offspring)
- Traits that affect an organisms schedule of reproduction and survival make up: LIFE HISTORY

- life history has 3 main variables
1) when reproduction begins (age at first reproduction or age at maturity)
2) how often the organism reproduces,
3) how many offspring are produced per reproductive episode
53.4 Life History Traits are products of Natural Selection

- Evolution and LIfe History Diversity
- Semelparity & Iteroparity
- Semelparity: "one-shot" pattern of big-bang reproduction, occurs in salmon, some plants such as agave.

- Iteroperity: opposite of semelparity. organisms produce relatively few, but large offspring, provisions are better (some lizards)

Evolution of Semelparity vs. Iteroparity
- 2 factors: (1) survival rate of offspring (2) likelihood adult will survive and reproduce again.
- survival rate of offspring and parent is low (unpredictable events) Semelparity is favored
- survival rate high, more dependable environment: Iteroparity is favored.
53.4 Life History Traits are products of Natural Selection

Trade-offs and LIfe Histories
- Plants and animals whose young are subject to high mortality rates often produce large numbers of relatively small offspring.
- Extra investment from parents increases offsprings chance of survival (Wal-nut & brazilian nut trees)

K-Selection:
- Selection for traits sensitive to pop density and are favored at high densities. Density-dependent selection
- lots of competition

r-Selection:
- selection for traits that maximize reproductive success in uncrowded environments (low density). Density-independent selection
- little competition
53.5 Factors that regulate pop. growth are density dependent

- Population Change and Pop. Density
- Density Independent & Density Dependent
- Density Independent: birth rate or death rate that does NOT change w/ pop density.

- Density Dependent: death rate that rises as pop density rises, as is birth rate that falls w/ rising density
53.5 Factors that regulate pop. growth are density dependent

Mechanisms of Density-dependent pop regulation
- Density-dependent regulation provides feedback regulation
- negative feed back between pop density and rates of birth and death, halts pop growth through mechanisms that reduce birth rates, increase death rates.
53.5 Factors that regulate pop. growth are density dependent

- Mechanisms of Density-Dependent Regulation
1) Competition for resources
2) Predation
3) Toxic Wastes
4) Intrinsic Factors
5) Territoriality
6) Disease
53.5 Factors that regulate pop. growth are density dependent

Population Dynamics
long term data showing fluctuation from year to year or place to place
- focuses on complex interactions between abiotic and biotic factors
53.5 Factors that regulate pop. growth are density dependent

Pop. Dynamics: Stability and function
- Dramatic pop changes can be multifactorial: good/bad weather, disease, food availability, predation.
53.5 Factors that regulate pop. growth are density dependent

Pop. Dynamics: Pop Cycles - Scientific Inquiry
- fluctuations in pop in cycles (hares: 10 year cycle)
- hypotheses?
1) cycles may be caused by food shortage in winter
2) predator-prey interactions - more than one animal other than lynx hunt the hare
3) Sunspot activity - undergoes cyclic changes.
53.5 Factors that regulate pop. growth are density dependent

Pop. Dynamics: immigration, emigration, and metapopulations
- immigration and emigration influence pop.
- pop gets crowded, resource competition increases, emigration increases
- immigration and emigration important when a number of local pop. are linked forming a metapopulation.

- Local pop in a metapop, occupying discrete patches of suitable habitat
- vary in size, quality
- patches w/ many individuals can supply more emigrants to other patches.
- if pop becomes extinct, patch occupied can be recolonized by immigrants from another pop.
53.6 Human Population

- Global Human pop
- pop increased slowly until 1650: 500 mill people
- Doubled to 1 bill w/in next 2 centuries
- 1930: 2 bill
- doubled by 1975 - more than 4 bill
- pop is now 6.8 bill
- increasing 79 mill each year
- growing at 200,000 each day
- end of 2050 about 7.8-10.8 bill
53.6 Human Population

Regional Patterns of Pop. Change
- Stable regional pop, birth rate = death rate

0 pop growth = high birth rate - high death rate
0 pop growth = low birth rate - low death rate

Demographic Transition:
53.6 Human Population

Age Structure
- Demographic variable is age structure
- relative number of individuals of each age in the population
- commonly graphed as "pyramids"
- percent on X axis, age on Y axis
53.6 Human Population

Estimates of Carrying Capacity
- First known estimate of pop capacity was 13.4 Billion, Anton Van Leeuwenhoek, discovered protists
- current researchers use curves to predict future max human pop
- others generalize max pop density X area of habitable land
- others use limiting factors: food
53.6 Human Population

Limits on Human Pop. Size
- human needs: food, water, fuel, shelter
- Ecological footprint summarizes aggregate land and water area req'd by each person, city, etc
- way to estimate: add up all ecologically productive land on planet, divide by pop
= 2 hectares (ha) per person
1 ha = 2.47 acres.