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132 Cards in this Set

  • Front
  • Back
Genetic Divergence what is it? How does it occur? What counters it?
Gradual accumulation of differences in the gene pools of populations
Natural selection, genetic drift, and mutation can contribute to divergence
Gene flow counters divergence
Morphology and Species the effects and things that it can't effect
Morphological traits may not be useful in distinguishing species
Members of same species may appear different because of environmental conditions
Morphology can vary with age and sex
Different species can appear identical
What is a species?
Species are groups of interbreeding natural populations that are reproductively isolated from other such groups.
The 7 isolating mechanisms are:
mechanical isolation, temporal isolation, behavioral isolation, ecological isolation, gamete mortality, hybrid inviability, and hybrid sterility
mechanical isolation
In mechanical isolation individuals cannot mate or pollinate because of physical incompatibilities
temporal isolation
individuals of different species reproduce at different times
behavioral isolation
individuals of different species ignore or don’t get the required cues for sex
ecological isolation
individuals of different species live in different places and never meet up
gamete mortality
gametes of different species are incompatible so no fertilization occurs
hybrid inviability
hybrid embryos die early or the new individuals die before they can reproduce
hybrid sterility
because hybrid individuals cannot make functional gametes
What are the 3 methods of speciation?
Allopatric speciation
Sympatric speciation
Parapatric speciation
Allopatric speciation what happens?
Speciation in geographically isolated populations
Some sort of barrier arises and prevents gene flow
Effectiveness of barrier varies with species
Extensive Divergence Prevents Inbreeding how?
Species separated by geographic barriers will diverge genetically

If divergence is great enough it will prevent inbreeding even if the barrier later disappears
Archipelagos what are they and how do they effect speciation?
Island chains some distance from continents
Galapagos Islands
Hawaiian Islands
Colonization of islands followed by genetic divergence sets the stage for speciation
sympatric speciation
Species forms within the home range of the parent species
parapatric speciation
Adjacent populations evolve into distinct species while maintaining contact along a common border
Sympatric Speciation in African Cichlids
Studied fish species in two lakes
Species in each lake are most likely descended from single ancestor
No barriers within either lake
Some ecological separation but species in each lake breed in sympatry
speciation by polyploidy how does this cause speciation
Change in chromosome number (3n, 4n, etc.)
Offspring with altered chromosome number cannot breed with parent population
Common mechanism of speciation in flowering plants
hybrid sterility
because hybrid individuals cannot make functional gametes
2 types of Patterns of Change in a lineage
anagenesis
Cladogenesis
What are the 3 methods of speciation?
Allopatric speciation
Sympatric speciation
Parapatric speciation
Allopatric speciation what happens?
Speciation in geographically isolated populations
Some sort of barrier arises and prevents gene flow
Effectiveness of barrier varies with species
Extensive Divergence Prevents Inbreeding how?
Species separated by geographic barriers will diverge genetically

If divergence is great enough it will prevent inbreeding even if the barrier later disappears
Archipelagos what are they and how do they effect speciation?
Island chains some distance from continents
Galapagos Islands
Hawaiian Islands
Colonization of islands followed by genetic divergence sets the stage for speciation
sympatric speciation
Species forms within the home range of the parent species
parapatric speciation
Adjacent populations evolve into distinct species while maintaining contact along a common border
Sympatric Speciation in African Cichlids
Studied fish species in two lakes
Species in each lake are most likely descended from single ancestor
No barriers within either lake
Some ecological separation but species in each lake breed in sympatry
speciation by polyploidy how does this cause speciation
Change in chromosome number (3n, 4n, etc.)
Offspring with altered chromosome number cannot breed with parent population
Common mechanism of speciation in flowering plants
2 types of Patterns of Change in a lineage
anagenesis
Cladogenesis
Cladogenesis
Branching pattern
Lineage splits, isolated populations diverge
Anagenesis
No branching
Changes occur within single lineage
Gene flow throughout process
What is the gradual model? Is it a good model or not?
Speciation model in which species emerge through many small morphological changes that accumulate over a long time period

Fits well with evidence from certain lineages in fossil record
what is the Punctuation model? Is it a good model or not?
Speciation model in which most changes in morphology are compressed into brief period near onset of divergence

Supported by fossil evidence in some lineages
Adaptive radiation causes?
Burst of divergence
Single lineage gives rise to many new species
New species fill vacant adaptive zone
Adaptive zone is “way of life”
Extinction how have they affected us sp far?
Irrevocable loss of a species
Mass extinctions have played a major role in evolutionary history
Fossil record shows 20 or more large-scale extinctions
Reduced diversity is followed by adaptive radiation
who survives an extinction?
Species survival is to some extent random
Asteroids have repeatedly struck Earth, destroying many lineages
Changes in global temperature favor lineages that are widely distributed
Taxonomy
Field of biology concerned with identifying, naming, and classifying species
Somewhat subjective
Information about species can be interpreted differently
Binomial system what is it and who developed it?
Devised by Carl von Linne
Each species has a two-part Latin name
First part is generic
Second part is specific name
higher taxa
Kingdom
Phylum
Class
Order
Family
Phylogely what is it and what are its practical applications?
The scientific study of evolutionary relationships among species

Practical applications
Allows predictions about the needs or weaknesses of one species on the basis of its known relationship to another
Five-Kingdom Scheme
Monera
Protista
Fungi
Plantae
Animalia
Are we related? How so or why not?
All species are related by descent

Share genetic connections that extend back in time to the prototypical cell
genetic disconnect
end of the line for a species. Mass extinctions are catastrophic events in which major groups abruptly and simultaneously are lost
genetic divergence
basis of life's diverstiy, as brought about by adaptive shifts, branching, and radiations. Rates and times of change varied within and between lineages
Genetic persistence
the basis of unity of life. The biochemical and molecular basis of inheritance extends from the origin of first cells through all subsequent lines of descent.
The big bang theory
12-15 billion years ago all matter was compressed into a space the size of our sun
Sudden instantaneous distribution of matter and energy throughout the known universe
Archaeon and earlier... Origin of earth was? Crust was formed when? What kind of evolution was occuring? What kind of cells developed?
4,600 mya: Origin of Earth
4,600 - 3,800 mya
Formation of Earth’s crust, atmosphere
Chemical and molecular evolution
First cells (anaerobic bacteria)
Earth forms
About 4.6 and 4.5 billion years ago
Minerals and ice orbiting the sun started clumping together
Heavy metals moved to Earth’s interior, lighter ones floated to surface
Produced outer crust and inner mantle
why is earth ideal for life?
Smaller in diameter, gravity would not be great enough to hold onto atmosphere
Closer to sun, water would have evaporated
Farther from sun, water would have been locked up as ice
What was the first atmosphere made up of?
Hydrogen gas
Nitrogen
Carbon monoxide
Carbon dioxide
No gaseous oxygen
Origin of organic compounds
Amino acids, other organic compounds can form spontaneously under conditions like those on early Earth
Clay may have served as template for complex compounds
Compounds may have formed near hydrothermal vents
chemical evolution
Spontaneous formation of porphyrin rings from formaldehyde
Components of chlorophylls and cytochromes
Discuss RNA and it's origin and the origin of life
DNA is genetic material now
DNA-to-RNA-to-protein system is complicated
RNA may have been first genetic material
RNA can assemble spontaneously
How switch from RNA to DNA might have occurred is not known
Proto cells
Microscopic spheres of proteins or lipids can self assemble
Tiny sacs like cell membranes can form under laboratory conditions that simulate conditions in evaporating tidepools
Nanobes may resemble proto-cells
Proterzoic eon what happens in the formation of the earth?
Origin of photosynthetic Eubacteria
Noncyclic pathway first
Cyclic pathway next
Oxygen accumulates in atmosphere
Origin of aerobic respiration
The first cells
Originated in Archeon Eon
Were prokaryotic heterotrophs
Secured energy through anaerobic pathways
No oxygen present
Relied on glycolysis and fermentation
What are the advantages of organelles
Nuclear envelope may have helped to protect genes from competition with foreign DNA
ER channels may have protected vital proteins
Theory of endosymbiosis
Lynn Margulis
Mitochondria and chloroplasts are the descendents of free-living prokaryotic organisms
Prokaryotes were engulfed by early eukaryotes and became permanent internal symbionts
Community
All the populations that live together in a habitat

Type of habitat shapes a community’s structure
6 factors that shape community structure
Climate and topography
Available foods and resources
Adaptations of species in community
Species interactions
Arrival and disappearance of species
Physical disturbances
Niche
Sum of activities and relationships in which a species engages to secure and use resources necessary for survival and reproduction
Realized and Fundamental niches
Fundamental niche
Theoretical niche occupied in the absence of any competing species
Realized niche
Niche a species actually occupies
Realized niche is some fraction of the fundamental niche
Species interactions discuss commensalism and mutualism
Most interactions are neutral; have no effect on either species
Commensalism helps one species and has no effect on the other
Mutualism helps both species
species interactions discuss interspecific competition and predation and parasitism
Interspecific competition has a negative effect on both species
Predation and parasitism both benefit one species at a cost to another
symbiosis
Living together for at least some part of the life cycle
Commensalism, mutualism, and parasitism are forms of symbiosis
Mutualism
Both species benefit
Some are obligatory; partners depend upon each other
Yucca plants and yucca moth
Mycorrhizal fungi and plants
yucca and yucca moth
Example of an obligatory mutualism
Each species of yucca is pollinated only by one species of moth
Moth larvae can grow only in that one species of yucca
mycorrhyzia
Obligatory mutualism between fungus and plant root
Fungus supplies mineral ions to root
Root supplies sugars to fungus
3 types of comeptition
Interspecific - between species
Intraspecific - between members of the same species
Intraspecific competition is most intense
forms of competition
Competitors may have equal access to a resource; compete to exploit resource more effectively
One competitor may be able to control access to a resource, to exclude others
competitive exclusion principle
When two species compete for identical resources, one will be more successful and will eventually eliminate the other
Hairston's experiment
Two salamanders species overlap in parts of their ranges
Removed one species or the other in test plots
Control plots unaltered
5 years later, salamander populations were growing in test plot
resource partitioning
Apparent competitors may have slightly different niches
May use resources in a different way or time
Minimizes competition and allows coexistence
predation
Predators are animals that feed on other living organisms
Predators are free-living; they do not take up residence on their prey
coevolution
Joint evolution of two or more species that exert selection pressure on each other as an outcome of close ecological interaction
As snail shells have thickened, claws of snail-eating crabs have become more massive
3 types of predator prey models and the different appearances of each
Type I model: Each individual predator will consume a constant number of prey individuals over time

Type II model: Consumption of prey by each predator increases, but not as fast as increases in prey density

Type III model: Predator response is lowest when prey density is lowest
variation in prey cycles
An association in predator and prey abundance does not always indicate a cause and effect relationship
Variations in food supply and additional predators may also influence changes in prey abundance
4 prey defenses
Camouflage
Warning coloration
Mimicry
Moment-of-truth defenses
predator responses
Any adaptation that protects prey may select for predators that can overcome that adaptation
Prey adaptations include stealth, camouflage, and ways to avoid chemical repellents
parasitism
Parasites drain nutrients from their hosts and live on or in their bodies
Natural selection favors parasites that do not kill their host too quickly
4 kinds of parasites
Microparasites
Macroparasites
Social parasites
Parasitoids
parasitoids
Insect larvae live inside and consume all of the soft tissues of the host

Used as agents of biological control

Can act as selective pressure on host
ecological succession
Change in the composition of species over time
Classical model describes a predictable sequence with a stable climax community
2 types of succession and what each does
Primary succession - new environments
Secondary succession - communities were destroyed or displaced
pioneer species
Species that colonize barren habitats
Lichens, small plants with brief life cycles
Improve conditions for other species who then replace them
climax community
Stable array of species that persists relatively unchanged over time
Succession does not always move predictably toward a specific climax community; other stable communities may persist
cyclical changes
Cyclic, nondirectional changes also shape community structure
Tree falls cause local patchiness in tropical forests
Fires periodically destroy underbrush in sequoia forests
restoration ecology
Natural restoration of a damaged community can take a very long time
Active restoration is an attempt to reestablish biodiversity in an area
Ecologists are actively working to restore reefs, grasslands, and wetlands
community instability
Disturbances can cause a community to change in ways that persist even if the change is reversed
Keystone species
A species that can dictate community structure
Removal of a keystone species can cause drastic changes in a community; can increase or decrease diversity
species introductions
Introduction of a nonindigenous species can decimate a community
No natural enemies or controls
Can outcompete native species
exotic species
Species that has left its home range and become established elsewhere
Becomes part of its new community
Can have beneficial, neutral, or harmful effects on a community
endangered species
A species that is extremely vulnerable to extinction
Close to 70 percent of endangered species have been negatively affected by exotic competitors
rabbits in austraillia
Rabbits were introduced for food and hunting
Without predators, their numbers soared
Attempts at control using fences or viruses have thus far been unsuccessful
where in the world is more diversity and where is les?
Diversity of most groups is greatest in tropics; declines toward poles
why are tropical species rich?
Resources are plentiful and reliable
Species diversity is self-reinforcing
Rates of speciation are highest in the tropics
distance effect on speciation
The farther an island is from a mainland, the fewer species
Closer islands receive more immigrants
Species that reach islands far from mainland are adapted for long-distance dispersal and can move on
area effect
Larger islands tend to support more species than smaller islands
More habitats
Bigger targets
Larger populations decrease extinction risks
Ecosystem
An association of organisms and their physical environment, interconnected by ongoing flow of energy and a cycling of materials
Autotrophs modes of nutrition
Capture sunlight or chemical energy
Producers
Heterotrophs Modes of nutrition
Extract energy from other organisms or organic wastes
Consumers, decomposers, detritivores
Consumer levels
Herbivores
Carnivores
Parasites
Omnivores
Decomposers
Detritivores
Trophic levels
All the organisms at a trophic level are the same number of steps away from the energy input into the system
Producers are closest to the energy input and are the first trophic level
Food Chain
A straight-line sequence of who eats whom
Simple food chains are rare in nature
Energy losses
Energy transfers are never 100 percent efficient
Some energy is lost at each step
Limits the number of trophic levels in an ecosystem
Biological magnification
A nondegradable or slowly degradable substance becomes more and more concentrated in the tissues of organisms at higher trophic levels of a food web
DDT and its effects
Synthetic pesticide banned in United States since the 1970s

Birds that are carnivores accumulate DDT in their tissues, produce brittle egg shells
Primary productivity Gross and net
Gross primary productivity is ecosystem’s total rate of photosynthesis
Net primary productivity is rate at which producers store energy in tissues in excess of their aerobic respiration
primary productivity varies how and why?
Seasonal variation
Variation by habitat
The harsher the environment, the slower plant growth, the lower the primary productivity
Ecological pyramids
Primary producers form a base
Successive tiers of consumer above them
Can be used to visualize:
Biomass
Dryweight of all the ecosystems organisms at each tier
Energy
Illustrates the transfer of energy through an ecosystem
Sunlight is captured at base and declines through successive levels
What happens to energy as it is used?
At each trophic level, the bulk of the energy received from the previous level is used in metabolism
This energy is released as heat energy and lost to the ecosystem
Eventually all energy is released as heat
biogeochemical cycle
The flow of a nutrient from the environment to living organisms and back to the environment
Main reservoir for the nutrient is in the environment
3 types of biogeochemical cycles and what each deals with
Hydrologic cycle
Water
Atmospheric cycles
Nitrogen and carbon
Sedimentary cycles
Phosphorus and sulfur
watersheds
Any region where precipitation flows into a single stream or river
Water seeps into soil or joins surface runoff
Water moves nutrients into and out of the surrounding ecosystem
Hubbard Brook Experiment
A watershed was experimentally stripped of vegetation
All surface water draining from watershed was measured
Removal of vegetation caused a six-fold increase in the calcium content of the runoff water
water use and scarcity
Most of Earth’s water is too salty for human consumption
Desalinization is expensive and requires large energy inputs
Irrigation of crops is the main use of freshwater
negative effects of irrigation
Salinization, mineral buildup in soil
Elevation of the water table and waterlogging
Depletion of aquifers
Carbon cycle atmospheric cycle
Carbon moves through the atmosphere and food webs on its way to and from the ocean, sediments, and rocks
Sediments and rocks are the main reservoir
Carbon in atmosphere what kind? How is it added? How is it removed?
Atmospheric carbon is mainly carbon dioxide
Carbon dioxide is added to atmosphere
Aerobic respiration, volcanic action, burning fossil fuels
Removed by photosynthesis
carbon in the ocean
Most carbon in the ocean is dissolved carbonate and bicarbonate
Ocean currents carry dissolved carbon
Greenhouse gasses
Greenhouse gases impede the escape of heat from Earth’s surface
Global Warming
Long-term increase in the temperature of Earth’s lower atmosphere
Carbon Dioxide increase
Carbon dioxide levels fluctuate seasonally
The average level is steadily increasing
Burning of fossil fuels and deforestation are contributing to the increase
Other greenhouse gasses and where they come from
CFCs - synthetic gases used in plastics and in refrigeration
Methane - produced by termites and bacteria
Nitrous oxide - released by bacteria, fertilizers, and animal wastes
Nitrogen cycle nitrogen atmospheric cycle
Nitrogen is used in amino acids and nucleic acids
Main reservoir is nitrogen gas in the atmosphere
Nitrogen fixation
Plants cannot use nitrogen gas
Nitrogen-fixing bacteria convert nitrogen gas into ammonia (NH3)
Ammonia and ammonium can be taken up by plants
Ammonification and nitrification how it occurs
Bacteria and fungi carry out ammonification
conversion of nitrogenous wastes to ammonia
Nitrifying bacteria convert ammonium to nitrites and nitrates
Nitrogen loss
Nitrogen is often a limiting factor in ecosystems
Nitrogen is lost from soils via leaching and runoff
Denitrifying bacteria convert nitrates and nitrites to nitrogen gas
How humans increase and decrease the amount of nitrogen in the air
Humans increase rate of nitrogen loss by clearing forests and grasslands
Humans increase nitrogen in water and air by using fertilizers and by burning fossil fuels
Too much or too little nitrogen can compromise plant health
Phosphorous cycle or sedimentary cycle
Phosphorus is part of phospholipids and all nucleotides
It is the most prevalent limiting factor in ecosystems
Main reservoir is Earth’s crust; no gaseous phase
human effects on phosphorus
In tropical countries, clearing lands for agriculture may deplete phosphorus-poor soils
In developed countries, phosphorus runoff is causing eutrophication of waterways