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52 Cards in this Set

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fossil
hard evidence of the existence of a species in the past
after death, bones can be destroyed by several forces
physical
chemical
biological (scavengers crunching up bones)
fossilization is promoted or enhanced if the animal dies:
in a cave
or near water
fossil record is non-random (i.e. biased):
1. where fossilization is likely to occur
2. which species are more likely to be fossilized
3. discovery of fossil bearing deposits
geological time scale
precambrian eon 4.6 bya - 570 mya
phanerozoic eon 570 mya - present
>>paleozoic era 570 mya - 225mya
>>mesozoic era 225 mya - 65 mya
>>cenozoic era 65 mya - present
cenozoic era epochs
paleocene: 65-55 mya
eocene: 55-33 mya
oligocene: 33-23 mya
(definitive proof of anthropoid ancestors of later monkeys/apes)
miocene: 23-5 mya
(definitive definition of apes evolve)
pliocene: 5-1.8 mya
pleistocene: 1.8 mya - 10,000 ya
holocene/recent: 10,000 to present
relative dating methods
stratigraphy
biostratigraphy
paleomagnetism
fluorine analysis
stratigraphy
dating of a specimen based on its position in the ground

based on law of superposition: upper sedimentary layers are younger in age than lower layers
biostratigraphy
faunal correlation

dating based on associated species whose evolutionary history is well known
paleomagnetism
dating based on shifts in the earth's polarity
fluorine analysis
dating of specimen based on relative amount of fluorine

if there is fluorine in the ground water, it can harden the bones. if 2 organisms die at different times, the bones that were younger will have more fluorine in them.

this aging technique can only be done at one geographical location
chronometric (absolute) dating methods
elements have isotopes
isotopes release radioactive energy to achieve a more stable state. The rate of radioactive discharged is the half-life. half-life is the amount of time it takes for one-half of radioactivity to be released

potassium argon dating. half-life is 1.25 billion years

carbon 14. half life is 5,730 years
fraudulent piltdown specimen
charles dawson discovered

it had different fluorine content compared to contemporaneous specimens
mandible had different fluorine content than cranium
bones were painted
mandible belonged to orangutan, teeth filed down. cranium was human.
suspects in piltdown hoax
charles dawson
arthur conan doyle
martin hinton
plate tectonic timeline
pangea: 224 mya
laurasia, gondwonaland: 135 mya
continents today: 65 mya

o North America and Europe were connected to one another during Paleocene and Eocene. Helps to explain geographical distribution of early primates.
o During the Cenozoic Era, Africa was an island continent prior to 16 million years ago. At about 16 million years ago, Africa contacted Southwest Asia. Helps to explain geographical distribution of apes during the Miocene.
how did lemurs colonize madagascar?
Best theory is that lemurs colonized Madagascar by crossing the Mozambique Channel on floating of masses vegetation.
how did monkeys colonize south america?
South America was an island continent long before the emergence of primates, and the Isthmus of Panama connected North and South America only about 2 million years ago.

Best theory is that the ancestor of New World monkeys colonized South America by crossing either the Caribbean Sea or Atlantic Ocean on floating masses of vegetation.
Paleocene (65-55 mya)
putative fossil primates found principally in north america and europe

paleocene species were not primates
-longer snout
-eyes not at front of skull
-no orbital bar
-smaller brain
-specialized dentition
-claws, not nails
-non prehensile hands/tail/feet
anatomies that suggest paleocene species may be primates
anatomy of molar teeth
middle ear
eocene (55-33 mya)
-first primates
-eyes face forward
-post orbital bar
-short snout
-prehensile hands and feet
-nails
-large cranial capacity (compared to paleocene species)
where fossils of eocene primates are found
found in north america and europe. also found in north africa and asia near the end.

north america and europe geographically connected in eocene by greenland.
environmental context in which eocene primates are found
tropical and subtropical forests
two lineages of eocene primates
Adapids may be ancestral to lemurs and lorises
Omomyids may be ancestral to tarsiers and anthropoids
inferences about eocene primates
-Some species were diurnal, whereas others were nocturnal
-Locomotion: quadrupedality and leaping
-Diet: insectivory; Frugivory-folivory
-Observation: absence of tooth comb
where fossils are found in Oligocene and dates of fossils
Site – Fayum, Egypt
Dates – 33 to 35 million years ago
environment of fayum site during ogliocene
tropical rainforest
taxonomy of oligocene anthropoids
Propliopithecines
-E.g., Aegyptopithecus
-Dental formula: 2, 1, 2, 3/2, 1, 2, 3
-Same dental formula as modern catarrhines (i.e., Old World monkeys, apes and humans)

Parapithecines
-E.g., Apidium
-Dental formula: 2, 1, 3, 3/2, 1, 3, 3
-Three premolar teeth in each quadrant of mouth; same dental formula as some modern platyrrhines (i.e., New World monkeys)
phylogeny of oligocene anthropoids based on dental formulas
Propliopithecines may be ancestor of catarrhines
-Evolutionary divergence between Old World monkeys and hominoids had not yet occurred

Parapithecines may be ancestor of New World monkeys
Inferences about Oligocene anthropoids: locomotion
Aegyptopithecus – quadrupedal, slow climber
Apidium – quadrupedal runner and leaper
Inferences about Oligocene anthropoids: body size and diet
Body size: 2 to 4 pounds for Apidium; 12 to 18 pounds for Aegyptopithecus

Diet: Frugivory
Inferences about Oligocene anthropoids: time of activity
diurnal
Inferences about Oligocene anthropoids: social organization
Social organization for Aegyptopithecus may have been polygyny

Sexual dimorphism in canine tooth size implies polygyny
sivapithecus
ancestor of orangutans. fossil specimen: GSP 15,000 (geological survey of pakistan)

chimps have broader septum than orangutans
miocene importance
earliest hominoids. based on y-5 cusp pattern of molar teeth.
where miocene fossils are found
africa in early & late middle miocene.

asia- middle and late micene
geological event that explains distribution of hominoids
-In the early Miocene, the Tethys Sea separated Africa from Europe and Asia.
-Approximately 16 million years ago, the Arabian plate partially separated from continental Africa and contacted southwest Asia.
-This established a migratory corridor.
-Subsequent to 16 million years ago, apes are found in Africa, Europe, and Asia.
grouping of hominoids based on geography
african hominoids (23 mya)
-ex: proconcil & kenyapithecus

european hominoids (16-11 mya)
-ex: dryopithecus-found in france, spain, italy, and greece

asian hominoids (15-7 mya)
-ex: sivapithecus - found in turkey, pakistan, india, and china
alternative grouping of hominoids based on dentition
proconsulines
-proconsul & dryopithecus
-compared to sivapitchecenes, smaller molars w/ thinner enamel and a thin, gracile mandible

sivapithecenes
-sivapithecus & kenyapithecus
-compared with proconsolines, have larger molars w/ thicker enamel and a thick, heavily buttressed mandible
implication of dental differences
-proconsulines and sivapithicenes had different diets.
-proconsulines frugivorous
-sivapithecenes- fruits, nuts, other hard objects

-dental differences associated with climactic changes during miocene

-during early miocene, broad tracts of forest. by middle and late miocene, the environment was more mosaic (forest, woodland, savana)
phylogenies
-sivapithecus may be ancestral to pongo (facial similarity b/w pongo and gsp 15,000)

-no direct ancestor has been identified for pan and gorilla
inferences
-proconsule has a monkey-like post cranium
-dryopithecus has anatomies suggsting arm-swinging locomotion
-sivapithecus has some monkey-like anatomies. implies independent evolition of some anatomies b/w pongo and pan/gorilla
gigantopithecus
-dates: 8-6 mya and as recently as 500,000 ya
-found in asia (pakistan, india, china, vietnam)
-anatomy: small incisors ,small canines, and huge molars with thick enamel and heavily buttressed mandible
-inferences: diet of tough consistency, body size 400-700 lbs, terrestrially adapted
early hominid evolution
humans are unique in locomotion among living mammals
which species is earliest hominid?
two traits to ID hominid:
bipedality
reduced canine size
period of hominid fossil sites
late miocene and early pliocene are in africa
east and north central africa
open air sites
dated by potasium argon, paleomagnetism, and biostratigraphy
famous sites:
-hadar, etheopia
-lake turkana, kenya
-olduvai gorge, tanzania
republic of south africa
limestone caves
dated by stratigraphy
famous site: taung
famous fossils
AL288 (lucy)
fossilized footprints - laetolia, tanzania
taung child
KNM-ER 1470 Bartlett homo
AL 288 (lucy)
-member of species Australopithecus afarensis
-dated to 3.2 mya
-found in Ethiopia
-40% of skeleton preserved (3.5 ft, 60 lbs)
member of species which was highly sexual dimorphic in body size.
fossilized footprints
-3.6 mya
-early evidence of bipedality
-australopithecus afarensis
Taung child
first australopithecene found (1924 by Raymond Darte)
-australopithecus africanus
oldowan
earliest stone tools
2.6 mya in Ethiopia also called pebble choppers.

australopithecus garhi may have been earliest stone user