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77 Cards in this Set

  • Front
  • Back

What is adaptation?

-Adaptation can mean a process or a state


-¨The molding by natural selection of a character to perform a function.¨


-Character can be adapted if there is evidence that it evolved for a function/purpose

What does function of an organism not explain?

-The current function does not equal the reason for existance


-Function can change throughout time

What are two examples of an organism having a character that was created for a different function than what it is currently used for?

-The ancestors of birds did not evolve feathers to fly, but to regulate body heat


-Giraffes do not have long necks for reaching higher leaves. Original reason is uncertain.

What is the adaptationist programme?

-The adaptationist programme is a style of thinking among evolutionary biologists

-Assumes near omnipotence of natural selection


-Natural selection forges organic design and fashion the best of all possible worlds

What does adaptationist programme do in practice?

-The adaptationist programme separates an organism, and find the adaptive story about each part


-They assume that trade-offs among competing body parts is the only constraint on perfection

What are some examples of evolutionary imperfection resulting from trade-offs?

-Giraffe females incidently obtaining long necks from male genes


-Hyena females have pseudo penises which can complicate delivery of pups


-Both the Brown Skua and hyena infant females commit siblicide (females having higher levels of testosterone).

What are some alternatives to adaptation?

-Darcy Wentworth Thompson proposed to use mathematics to describe the shapes, chemistry and physics of organisms (allometric growth)

What is allometric growth?

-Allometric growth is the proportioning that helps give its body its specific form


-Many similar species have the exact same allometric growth, which can be explained mathematically

What is paedomorphosis?

-Paedomorphosis is when the rate of reproductive development accelerates compared to somatic development


-This means sexually active species retain juvenile phenotype, such as axolotl

How does pattern formation effect plant and animal mechanisms.

-Pattern formation in plants and animals have similar spatial organisation of tissues and organs


-These mechanisms explain how distantly related organisms can have similar anatomical arrangement

How can pattern formation be controlled?

-Through homeotic genes


-Drosophila larvae can have the undeveloped ´leg cells´ taken from one and translated into another


-The translantee will grow with an extra developed pair of legs


-This can be done with a number of body parts

What are hox genes?

-Hometic genes that provide information on the development of fins in fish and limbs in tetrapods

What are the three types of genetic markers?

1) Chromosomes - the banding pattern on chromosomes is the most commonly used marker


2) Protein polymorphs - electrophoretically detect detectable differences in enzymes are also widely used genetic markers.


3) Microsatellite (nuclear)/Mitochondrial DNA - recently developed and widely used

What are the uses of genetic markers?

-Phylogeny reconstruction


-In determination of levels of variation of a pop


-To estimate the effects of genetic drift/bottlenecks


-To determine levels of relatedness


-To detect species

What is minisatellite DNA?

Digested DNA placed on strips, able to compare between parents and offspring.

What is the meaning of species?

-In latin ´kind´ or ´appereance´


-To determine this, scientists compare morphology, physiology and biochemistry

What is the biological species concept?

-The biological species concept states that a species is a group of pops whose members have the potential to interbred and have fertile offspring


-Do not breed successfully with other pops


-Gene flow between populations holds the phenotype of a pop together

What is reproductive isolation?

-Reproductive isolation is the existence of biological factors that impede two species from producing viable, fertile offspring.


-Can be classified by whether factors act before or after fertilisation.

What are hybrids?

-Hybrids are the offsprings of two different species

What are the prezygotic barriers of reproductive isolation?

-Prezygotic barriers block fertilisation from occuring by:


-Impeding different species from attempting to mate


-Hindering fertilisation if mating is successful

What are the habitat isolation barriers of reproductive isolation?

-Habitat isolation is when two species encounter each other rarely, because they occupy different habitats, even though they are not separated by physical barriers.

What are the temporal isolation barriers of reproductive isolation?

-Temporal isolation occurs when species that breed at different times of day/year/seasons cannot mix their gametes.

What is behavioural isolation barriers?

Behavioural isolation means courtship rituals of one species is unique, and creates a barrier for other species from mating with them (since they cannot perform the ritual).

What is mechanical isolation?

-Mechanical isolation is when morphological differences (physical features) prevent successful mating


-An example is with snails who have certain shell orientations, stops successful mating.

What is gametic isolation?

The sperm of one species may not be able to fertilise eggs of another species.

What is postzygotic barriers of reproduction isolation?

Postzygotic barriers prevent the hybrid zygote from developing into a viable, fertile adult:


-Reduced hybrid viability/fertility


-Hybrid breakdown

What does reduced hybrid viability refer to?

-Reduced hybrid viability means the genes of two different parent species may interact and impair the hybrids development


-Hybrids may also be sterile

What is hybrid breakdown?

-Hybrid breakdown is when first-generation are fertile, but when they mate with another species (hybrid or parent), the offspring of the next generation is feeble/sterile

What are some limitations of the biological species concept?

-Many species are physically or ecologically distant but can interbred (grizzly/polar bear hybrids)


-The biological species concept cannot be applied to fossils


-Can not be applied to asexual organisms (and all prokaryotes)

What is the morphological species concept?

-Morphological species concept can be applied to both sexual and asexual species (unlike reproductive isolation theory)


-Relies on subjective critieria


-Defines a species by structural features


-Requires understanding in morphology to identify homologies, sexual dimorphic organisms, or organisms with complex life cycles

What is the ecological species concept?

-The ecological species concept views a species in terms of its ecological niche


-Applies to sexual/asexual species and an emphasis on disruptive selection

What is the phylogenetic genetic species concept?

-The phylogenetic species concept defines a species as the smallest group of individuals on a phylogenetic tree


-Emphasis on descent


-Applies to sexual/asexual


-No clear threshold to what a new species is


-Can split species

What is the specific mate recognition system?

-The specific mate recognition system defines a species as a group of organisms that recognise each other as mates


-´strucuturally coupled´


-emphasis is on behaviour and role of behaviour in fertilisation

What are the properties of the mate recognition system?

-The mate recognition is mediated by hormones and triggered by external factors


-Results in efficient fertilisation


-Male/female system operates within a content of various environmental factors


-This system is generally stable except during speciation

What is speciation and in what ways does it occur?

-Speciation can take place with or without geographic seperation


-Can occur in two ways:


-Allopatric speciation


-Sympatric speciation

What is allopatric speciation?

-Allopatric speciation is when gene flow between populations is interrupted


-When a population is geographically divided into geographically isolated subpops


-They evolve individually


-If reintroduced, they cannot interbreed

What are some properties of allopatric speciation?

-Seperated pops may evolve independently through mutation, NS and genetic drift


-Barriers to reproduction are intrinsic, separation is not a biological barrier

What is evidence of allopatric speciation?

-Regions with many geographic barriers have more species


-Reproductive isolation between populations generally increases as the distance between them increases

What are the two types of speciation?

Allotropic speciation


Sympatric speciation

Sympatric matings - Allopatric matings


(AmAf + Bm+Bf) - (AmBf + AfBm)


A = population 1


B = Population 2


What value is complete isolation between two populations?


What value is non-assortive mating?

-If a population/gender has a value of 1, it means mating is occurring.


-Complete isolation is:


(1 + 1) - (0 + 0) = 2


-Non-assortive mating is:


(1 + 1) - (1 + 1) = 0

What is sympatric speciation?

Sympatric speciation is when speciation takes place in geographically overlapping areas

What is polyploidy?

-Polyploidy is the presence of extra sets of chromosomes due to accidents during cell division

What is an autopolyploid?

-An autopolyploid is an individual with more than two chromosomes sets (from one species)

What is habitat differentiation?

-A form of sympatric speciation


-Can result in new ecological niches


-An example, an animal population eats one plant. A new plant is introduced into the ecosystem, and half the population begins eating that. After enough generations, they cannot eat the others fruit and only reproduce amongst their subpopulation.

What is sexual selection?

-A form of sympatric speciation


-Sexual selection can drive sympatic speciation


-sexual selection for mates of a different colour has contributed to speciation of cichlid fish in lake victoria

What is anallopolyploid?

-An allopolyploid is a species with multiple sets of chromosomes derived from different species (may only be able to reproduce with other allopolyploids)

What is hybridisation?

When two sister species are reintroduced and produce offspring?

What is a hybrid zone?

The area in which members of different species mate and produce hybrids

What are the three possible outcomes when species meet in a hybrid zone?

-The strengthening of reproductive barriers


-The weakening of reproductive barriers


-The continued formation of hybrid individuals

What process occurs when hybrids are less fit than parent species?

-Reinforcement of reproductive barriers stop hybrid offspring


-Separates parent species further


-Eventually stops hybrid offspring from being reproduced

What process occurs when hybrids are of equal fitness as parent species?

-Both parent populations coalesce


-Differentiation between parental populations decreases


-Substantial gene flow amongst populations


-If gene flow is great enough, the two species fuse into one species

What process occurs when hybrids are fitter than parental species?

-Reproductive barriers lower


-Formation of new hybrid species

At what rate can speciation occur?

-Can occur slowly or very rapidly


-4, 000 years (some cichlids)


-40, 000, 000 years (some beetles)


-6, 500, 00 years on average

How can the rate of speciation be determined?

-Patterns in speciation can be studied using the:


-fossil record


-morphological data


-molecular data

What relationship does macroevolution and speciation have?

Macroevolution is cumulative effect of speciation and extinction events

What is synthetic evolution?

-Synthetic evolution is the merging of Mendelian genetics and Weissman´s theory


-When these theories merged, it changed Darwinism to Synthetic Evolution

What is the Linnaean system of taxonomy?

-The linnaean system distinguishes species based on sameness and differences in morphology.


-This reinforces the notion of sameness (genus) and differences (species)

What is the Hierarchial Classification system of taxonomy?

-Linnaeus´ system led to Hierarchial Classification of species


-A system for grouping species in increasingly broad categories

When naming a species, what are the two criteria?

The name of the genus first before the species name, then underline or italicize the name.

What does a divergence mean in the evolutionary tree?

-The divergence of two species from a common ancestor


-The ¨Deeper¨ the points of separation, the greater the amounts of divergence.

What are the two currently used theories in taxonomy?

-The Phylogenetic Systematics


-Cladistics

What is Homology?

When two different species have individually evolved from a common ancestor.

What is Analogy?

-When two morphologically similar species have evolved from two separate common ancestors


-They are similar due to being driven by similar environmental factors

What is a Taxon?

A Taxon is a species at the top of a phylogenetic tree

What is a clade?

A clade is a group of taxons (species) at the top of a phylogenetic tree

What is a phylogenetic tree?

-A phylogenetic tree is a diagram like a tree.


-The branches represent the evolutionary lineages


-It shows the common descent of species in the higher taxa.

What is a cladogram?

-a ¨Nested hierarchy of taxa¨links several species of animals together based on morphological similarities


-Usually an outlier species outside the clade for comparison


-Ie Bass, Lizards, Horses and Monkeys all have vertebrae/jaws,


-Octopodes do not and are outliers

What can be inferred using a cladogram?

-Using a clade to determine the morphological chatacteristics of species, we can determine what the character the common ancestor might have had


-We can also determine what the common ancestor did not have from the outlier group in the cladogram

What is the difference from species at the top and at the bottom of a cladogram/phylogenetic trees?

-Species at the top of the tree are more derived

What makes a taxon monophyletic?

A taxon is monophyletic if it includes the most recent common ancestor, all members of that group, and the most derived descendants.




Common ancestor>members of group>all derived descendants (alive today)

What makes a taxon paraphylitic?

A taxon is paraphyletic if it includes the most recent common ancestor of all members of a group, but not all descendants of that ancestor.




Common ancestor>members of group>not all derived descendants

What makes a taxon polyphyletic?

A taxon is polyphyletic if it does not include most recently common ancestor of all members of a group




No common ancestor>not all members of group>not all derived descendants

What are the two principles of traditional evolutionary taxonomy?

1) Common descent


2) Amount of adaptive change


-Evolutionary taxa must have a single evolutionary origin and must show unique adaptive featuers

What did George Gaylord Simpson argue about the evolutionary tree?

-A higher taxon on the evolutionary tree should be given a higher status if it represents an adaptive zone

What are adaptive zones?

-Adaptive zones are places in the evolutionary tree that tends to have a faster rate of evolution, like the penguin branch in the bird tree


-Or humans in the great age tree

What are the differences of identifying species in biology and taxonomy?

-In biology, a species is a self-defining natural group


-In taxonomy, a species is a taxon like all others in its genus/genus/family, etc

What is the use of barcoding?

-Barcoding allows us to compare DNA from two organisms and determine the level of difference (and figure out the level of speciation)