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46 Cards in this Set

  • Front
  • Back
Arthropoda
Impact on humans
- Good: pollination, food
- Bad: Competitors for crops, disease transmission, bites, stings, etc
Arthropoda
Basic morphology
- Metamerism (less conspicuous)
- Fused segments (Tagmata, usually 2 or 3)
- Jointed lateral appendages
- Thick exoskeleton
- Hemocoel present
Arthropoda
Exoskeleton
- Rigid plates w/ flexible areas
- Individual segment is a somites
- Plates/sclerites on dorsal, ventral and lateral
- Also contain; pigments, phenols and inorganic material.
- Resistant and flexible
- Soft when secreted, hardened by sclerotization and minerlization
Arthropoda
Epicuticle
- Outter epicuticle w/ outter cement layer(protect against microbial invasion), a wax layer and an inner cuticulin layer (proteinaceous with 2 layers).
- Wax layer resist desiccation
- Wax glands also secrete; wax bloom (extend from surface add color and absorp light)
- No chitin in epicuticle
Arthropoda
Procuticle
- Layer beneath epicuticle
- Thicker than epicuticle
- composed of protein and chitin (polysaccharide, simular to cellulose) Long chains of chitin form microfibrils.
Arthropoda
Sclerotization
- Major hardening of skeleton due to cross linkage between sclerotin protein molecules.
- "tanning"
- Exocuticle more sclerotization than endocuticle
Arthropoda
Mineralization/Calcification
- In crustaceans
- Mineral deposit
- Outter portion of procuticle calcified, inner portion not.
Arthropoda
Appendages
- Attached at inward projetin apodemes
- Jointed
- Each segment with ventrolateral pair
- Each portion of appendage is a podiet
- Derived from; parapodia of polychaete or Phylum Onychophora
- Either uniramous or biramous
- Includes mouthparts and locomotory.
- Serial homology.
Arthropoda
Musculature
- Striated muscle(rapid contraction and relaxation)short bands from one segment to the next.
- Extrinsic limb (inside body to inside appendage) and Intrinsic (one limb podite to another)
- Antagonistic pairs (flexor and extensor)
- Fulcrum point condyle (ball and socket)
Arthropoda
Nervous system
- Brain in 2 or 3 regions (seperate ganglion); Protocerebrum (anterior w/ optic), Deutocerebrum (antennal nerves), tritocerebrum (labium)
- supraoesophageal ganglion (above digestive tract) and suboesophageal ganglion (below digestive tract)
- Ventral nerve cord with segmental ganglia.
- Condensation : fusion of segmental ganglia.
- Graded response (not all or nothing)
Arthropoda
Digestive System
- "outside" the animal
- Specialization of regions;
Foregut: lined with cuticle, simular to crop, ingestion, transport, storage
Midgut - not line, absorption, enzyme production
Hindgut - lined, water absorption, fecal production
Arthropoda
Circulatory system
- Open system
- Tissue in hemolymph (intestinal fluid) with haemocyanin, haemoglobin, to transport nutrients, wastes, gases, and immune responses.
- heart
- Hemolymph enters heart through Ostia.
Arthropoda
Gas exchange
- Aquatic: large SA/V ratio, diffusion across body or gills present
- Terrestrial: internal gills (Book lungs), and tracheae.
Arthropoda
Excretion and Osmoregulation
- Tubular structures
- Closed on inside
- Glandular excretory structures (maxillary or antennal gland in crustaceans and coxal gland in arachnids)
- Malpighian tubules.
- Nitrogen wates as ammonia (aquatic species) and uric acid (terrestrial species)
Arthropoda
Visual System
- Ocelli (simple eyes) non image forming, detect light changes
- Compound eyes, image forming, composed of facetes and ommatidia (300 - 25,000per eye)
--> Fixed focus lense (cornea), crystalline cone, retinular cells (up to 8) and cylindrical cells (collars)
--> Pigments in Rhadomeres, with composite unit as rhabdom.
- Resolving power determined by; angle, blending of light, angle of adjacent ommatidia, no of ommatidia.
Arthropoda
Reproduction
- Dioceious, some sequential hermaphrodites, some parthenogenetic (unfertilized = male)
- Internal fertilization
- Determinate growth
Arthropoda
Ecdysis
- Periodic moulting of exoskeleton
- Instars: intermoult stages
- Under neural and hormonal control. Insects hormone is stimulated from brain, Crustaceans; hormone is inhibited by neurosecretory complex.
- Exuviae: shed skin
- Fixed number of moults in insects and unlimited in crustaceans.
Subphylum Trilobitomorpha
Apomorphy
- Body divided into three regions by two anterior-posterior furrows
Class Trilobita
Characteristics
- Dorsoventrally flattened
- 4,000 - 15000 spp. Benthic or pelagic,
- 3 Tagmata; cehalon, thorax and pygidium
- Compound eyes
- Biramous limb
- 1 Pair antennae
Subphylum Chelicerata
Apomorphy
- No antennae
- Chelicerae (first two appendages modified for feeding)
- Two tagmata (prosoma and opisthosoma)
Chelicerata
Appendages
- Pedipalps: 2nd pair, sensory preception, catching prey, conveying sperm.
- 4 pair walking legs
- Each have gnathobases (toothed edges of appendage)
Chelicerata
Classification
Spp #
3 classes
- Merostomata
- Pycnogonida
- Arachnida
--> 75,000 ssp.
Merostomata
Apomorphy
- Class w/in Chelicerata
- Book gills
- Telson (tail spike)
Merostomata
Classification
- Order Eurypterida: sea scorpion (extinct)largest body.
- Xiphasura : horseshoe crabs, 4ssp left, all marine, chelicerates w/ compound eyes
Class Pycnogonida
Characteristics
-Within subphylum Chelicerata
- Sea spiders
- 1,000 ssp.
- Marien, no tagmata
- Proboscis (suck fluid)
- Predators and parasites
- 4 -6 pairs walking legs, long.
- Males w/ ovigers to brood eggs
Class Arachnida
Characteristics
- W/in Subphylum chelicerata
- Spiders and mites
- Terrestrial
- 74,000 ssp.
- Prosoma w/ shield carapace, opisthosoma w/o appendages.
- Simple eyes (0-4)
- Book lungs and/or tracheae.
- Excretion via coxal glands/malpighian tubules
Class Arachnida
Apomorphy
- Single median genital opening.
Oder Araneae
Characteristics
- Spiders
- Spinnerets, silk glands
- Chelicerae modified into fangs
- all are predators
Order Acari
Characteristics
- Mites and ticks
- 1 million undescribed ssp.
- Many parasites
- Prosoma and opisthosoma fused
Arachnida
Other classes...
Order Solifugae : wind scorpions
Oder Uropygi: whip scorpions
Order Amblypygi: tailess whip scorpions
Opiliones: daddy long legs
Order Scorpiones: Scorpions
Subphylum Mandibulata
Classification
- Class Crustacea
--> Subclass Malacostraca
- Class myriapoda
- Class Inseta
Mandibulata
Apomorphy
- Mandibles (biting jaws)
Class Crustacea
Apomorphy
- pairs of antennae
- Nauplius larva (3 pr. appendages and single eye)
Subclass Malacostraca
Characteristics
- W/in crustacea
- 75% of crustaceans
- 1-3 thoracic limbs (Maxillipeds)
- Pereopods: thoracic limbs
- Pleopods: abdominal limbs (swimming)
- Uropods :final segment (paddle)
Order decapoda
Characteristics/Appendages
- Lobser, crayfish, shrimp, crab
- Pigment cells (Chromatophores)
-5 pair pereopods
Order Isopod
Characteristics
- 10,000 ssp
- Marine, FW and terrestrial
- Dorsoventrally flattened
- Uniramous limbs
Amphipods
Characteristic
- W/in crustacea, malaostraca
- 6000 ssp, flattened laterally
- Marine, FW, semi terrestrial
Myriapoda
Characteristics
- Centipedes and Millipedes
- Two tagmata Head + Trunk
- no compound eyes
- 1 pair antennae
- Simular segments
- Uniramous limbs
- Tracheae
- Malphighian tubules
Order Chilopoda
- 3000 Ssp
- Terrestiral
- Unwaxed cuticle and unclosable spiracles
- moist habitats/nocturnal
- Predators (poision fangs), mandible, 2 pr. maxillae
-1 pair legs per body segment
- Can be long leg (fast) or short leg (slow)
Order Diplopoda
- 10,000 ssp
- Terrestrial, detritivores, herbivores
- Diplosegments, 2 pr legs per segment
-- Mandible, 1 pr. maxillae, gnathochilarium
- Calcium hardened cuticle
- Defense secretions
Class Insecta
Characteristics
- 1 million ssp.
- 3 tagmata
- 3 pairs of legs
- 2 pr. wings
- 1 pr. antennae
- Compound eyes
- Mandibles, maxillae and labium
Insecta
Apomorphy
- Fusion of 2nd maxillae to form labium
- Loss of all abdominal appendages
Insecta
Pre-requisites for wings
1. Striated muscle (more responsive than smooth)
2. Hard skeleton (leverage)
3. Sclerotin (light-weight strong and flexible)
Two groups of Pterygota
- Major evolution in flight
--> Paleoptera: 8000 ssp. wings can NOT be folded flat onto back
--> Neoptera: 1 million ssp. Wings CAN be folded flat onto back.
Insecta
Wings
- Key innovation
- Thin outfoldings of body wall
- network of tracheae and blood vessels
Two theories of
origin of wings...
- Paranotal theory: dorsla plate projections (not from pre existing limb structures) For thermoregulation, gliding.
- Exite theory: articulated gill plates. for respiration and thermoregulation. Skimming, flapping. *** Has the most support.