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66 Cards in this Set

  • Front
  • Back
What is Sound, physically?
- changes in pressure over time, transmitted through a medium - air or water
Sources of Sound

- how is it produced
- initiated by movement that disturbs air molecules
- molecules collied w/ each other
- this results in changes in air pressure that spreads from the source
How Sound Propagates through the air

Tuning Fork example
- Tuning fork gets struck
- Tine moves out (right)> collides w/ air molecules to right. Compressing molecules in section close together.
- Tine moves in (more left than original)> Results in rarefaction, less than normal air pressure.
- Time moves back out> collides w/ air molecules again, and pushes those molecules out further colliding into further out air molecules.
Definition:

Soundwave
- Waves of pressure changes in air caused by vibrations of a source
Definition:

Cycle
In a soundwave, a repeating segment of air pressure changes.
Definition:

INverse Square Law
- Sound E on wave front decreases w/ distance from source.
- Falloff in Sound E w/ distance: Inverse Square Law
- Sound decreases in proportion to square of the distance from source
- Sounds at a distance are much quieter
3 Most Important physical dimensions of sound are:
1) Frequency: pitch
2) Amplitude: loudness
3) Waveform: timbre (sound quality)
What is a Periodic Sound Wave?
- Waves in which the cycles of compression and rarefaction repeat in a regular, or periodic, fashion.
What is a Pure Tone?
- simplest periodic sound wave
- sound wave which air pressure changes over time according to a mathematical formula called Sine Wave or Sinusoid
- can be produced by an electronic signal generator and can be approximated by a flute
Definition:

Frequency
The physical dimension of sound that is related to perceptual dimension of pitch
- expressed in hertz (eg. 1000 cycles/sec = 1000Hz)
- number of cycles per second of a periodic sound wave
What is PItch?
Perceptual dimension of sound that corresponds to physical dimension of frequency
- perceived highness or lowness of a sound.
> HIgh pitch: piccolo
> low pitch: tuba
What is the sound detection range for humans?
20-20,000 HZ
- outside range, is inaudible to humans (no matter how loud)
What is Amplitude?
- Difference between the Max and minimum sound pressure in a sound wave
- distance from peak to trough.
- perceptual dimension of loudness
Define:

Loudness
- Perceptual dimension of sound related to physical dimension of amplitude
- how intense or quiet a sound seems
- also depends on frequency of sound and other factors
What are Decibels (dB)?
Unit to measure sound amplitude; logarithmically related to sound pressure measure in micropascals.
- amplitude is measure in terms of sound pressure
What is:

dB SPL

dB SPL = 20 log (p/p0)
decibel sound pressure level

P0 is set at agreed-upon 20uPa, thus amplitude of a sound w/ pressure p = 20,000 uPa

20 log (20,000/20) = 20 log 1,000 = 20x3 = 60dB SPL
Calculating Decibels
- 20 dB = 10x increase in sound pressure
- 100 dB = 100,000x more intense

20=20 log (p1/p0) 100=20 log (p1/p0)
1= log (p1/p0) 5=log (p1/p0)
10=p1/p0 100,000=p1 / p0
Absolute Threshold for Hearing
- Absolute threshold is the intensity of least intense sound that can be heard.
- not a single value
- depends on sounds freq
Absolute Threshold for hearing

- experiment
- psychophysical experiments
- P sits in sound attenuating chamber, w/ high fidelity speaker 2m in front of head, at ear level
- chamber reduces outside noise
- walls absorb sound waves, to reduce echoes
- heartbeat is present
- amplitude of tones, measure when taking P out, and placing mic where head was.
Audibility Curve
Curve showing minimum amplitude at which sounds can be detected at each frequency
- audibility threshold at lower and higher freq that people can hear (20-20,000 hz) is greater that freq in middle range (500-5,000)
- auditory sensitivity is max in this middle range, range of freq present in most human speech sounds.
Waveform
- sounds produced by vibrations of objects such as vocal cords & clarinets - consist of multiple pure tones added together
Waveform

- Fourier Analysis
- Fourier Analysis: mathematical procedure for decomposing complex waveform into a collection of sine waves w/ various freq & amplitude

Eg) Fourier analysis shows complex waveform is sum of 3 sine waves.
- Amplitude of complex waveform at any given time is sum of amplitudes-positive or negative- of the component wave at that time.
Waveform

- Fourier Spectrum
A depiction of the amplitudes at all frequencies that make up a complex waveform
- each sine wave is represented as a vertical line, single freq w/ a given amplitude
Waveform

- Fundamental Frequency
- Frequency of the lowest-freq component of the complex waveform
- determines the perceived pitch of the sound.
Waveform

- Harmonic
- Each component of a fundamental spectrum is called a harmonic
- first harmonic is the fundamental freq, 2nd harmonic is next lowest-freq component, and so on.
- 2nd & higher harmonics also called: overtones

- 2nd & higher harmonics are integer multiples of the fundamental freq.

eg) fundamental freq of the note A880 produced by clarinet is 880Hz,
- 2nd harmonic is 880x2=1760Hz,
- 3rd harmonic is 880x3= 2640
Waveform

- what is Timber?
- Two complex sounds that have same pitch & loudness, but dont sound the same.
- difference in sound quality between two sounds
- for complex periodic sounds, timbre is due to dif in relative amplitudes of sounds overtones
- perceptual dimension of sounds is that is related to physical dimension of waveform
- low amplitude overtones contribute less to timbre than high-amp overtones
Differences in Timbre

Flute vs Violin
- Flute spectrum is dominated by first 2-3 harmonics, give flute a sound quality closer to that of a pure tone

- Violin has significant E at many of the higher harmonics, giving the violin a "richer" sound quality

- diff in relative amplitudes of harmonics are what give each sound its distinctive timbre.
What happens if only the fundamental freq is remove, without removing any other harmonics?
- There is a diff in sound quality
- pitch of the sound seems to be same as before, even though the fundamental freq, which determines the pitch of a complex periodic sound, isn't present.
- Called Illusion of Missing Fundamental
The Ear

- What is it, and what does it do?
- Peripheral part of auditory system
- Transduces sound into neural signals
- divided into: outer, middler, and inner ear
Outer Ear
3 parts
- Pinna, auditory canal, Outer surface of Tympanic Membrane
- Function: outer ear funnels sound onto tympanic membrane - which vibrates in response to sound waves. > vibes are transmitted into middle ear.
Outer Ear

- Pinna
Outermost portion of ear
- shaped like funnel to direct sound into ear canal
- aids in sound localization
Outer Ear

- Auditory Canal
- Narrow channel that funnels sound waves gathered by Pinna onto tympanic membrane

- amplifies freq in the range of 2000-5000 Hz
Outer Ear

- Tympanic membrane
aka; eardrum
- thin, elastic diaphragm forms seal between outer & mid ear
- vibrates in response to sound waves that strike it.
Middle Ear
- responsible for amplifying and transmitting sound signal between outside air (eardrum) and inner ear (oval window)
Middle Ear

- Ossicles
Three smallest bones in the body
- transmit sound E from tympanic membrane to inner ear
3 Bone:
- Malleus (or hammer), Incus (or anvil), Stapes (or stirrup)
- bones are connect to one another, malleus is connected to tympanic membrane> when tym. membrane pushes on the malleus, malleus pushes on incus, displaces stapes, pushes on Oval Window> sends vibration to cochlea (liquid chamber)
Middle Ear

- ossicle anatomical advantages
1) Tympanic membrane is 15-20x larger than Oval Window.
- sound E collected by tymp mem is concentrated on smaller area, amplifying effect!

2) Ossicles produce lever action
- small movement at Malleus (near axis point), causes large displacement of the stapes (moment)
- magnifies vibrations

> these two characteristics can amplify sounds in air by 20-30dB
Middle Ear

- Acoustic Reflex
- reflex contractions of 2 tiny muscles in ear
- responds to high intensity sounds
- dampens movements of ossicles, helps protect auditory system from damage due to loud noises

- Limited capacity, responds to lower freq.
- vulnerable to loud high-freq noise
- Also takes more than 1/10th of sec to occur, too slow for brief high-intensity sounds
Middle Ear

- Eustachian Tube
Tube connecting the mid ear and top part of throat; normally closed but can open briefly (yawning, swallowing) to equalize air pressure in mid ear w/ air pressure outside.

eg) airplane or tall elevator,
- Unequal air pressure when going up tall bldg
- air pressure outside tymp mem goes down
- air pressure in mid ear doesn't change
- greater pressure on inner ear, makes membrane stretch, dampens vibrations> causes muffling sound.
Inner Ear

- Cochlea
- Snail shaped compartment
- has 3 chambers: Vestibular canal, Cochlear duct, Tympanic Canal
- Reissner's membrane separates chambers
Inner Ear

- Cochlear duct & organ of Corti
- one of three chambers in cochlea
- resting on basilar membrane w/ in cochlear duct is Organ of Corti> responsible for auditory transduction
- Cochlear duct is filled w/ endolymph, fluid that facilitates auditory transduction
- Organ of Corti contains hair cell receptors
Inner Ear

- Vestibular & Tympanic Canals
- chambers of cochlea
- both filled w/ perilymph, fluid similar to CSF.
- perilymph carries vibrations through cochlea
Inner Ear

- Semicircular Canals
1) Vestibular system
- sense organs used to produce neural signals carrying info about balance and acceleration; includes semicircular canals and otolith organs

2) semicircular canals
- part of vestibular system; perpendicular hollow curved tubes in skull, filled w/ endolymph, responsible for signaling head rotation.
Organ of Corti
- w/in cochlear duct
- rests on basilar membrane
- responsible for auditory transduction
- 3 components: two sets of neurons:
- Inner Hair Cells and Outer Hair Cells, and Tectorial Membrane
Hair Cells
- One row of 3,500 inner hair cells
- 3 rows of 12,00 outer hair cells total

> Outer hair cells are cylindrical, stereocilia are attached to tectorial membrane, amplify and sharpen responses of inner hair cells, 5% of auditory nerve fibres

> Inner Hair cells: are pear shaped, tips of inner h.c stereocilia float free in endolymph, Inner h.c responsible for tranducing sound into neural signals, 95% of auditory nerve fibers
Tip Link open ions channels on
a) bending of hair cell stereocilia increases tnesion on the tip links connecting them
- tip links pulls open channel, positively charged Ca & K ions enter hair cell
What do outer hair cells do?
- depol of cell membrane results in change in shape of protein, called prestin in the membrane
- shape change causes cell body and stereocilia to execute physical movements similar to stretching & contracting - motile response

> changes in length of outer H.C, magnify movements of basilar membrane in regions w/ characteristics freq corresponding to freq in the sound> means inner HC in those regoins send stronger signals in response to sound> motile response amplifies sound

> Magnified movements ocur in narrow regions of bas membrane, means iiner HC send more signals that are more freq specific - motile response sharpens response to freq in sound
Basilar Mmebrane & Neural encoding of pitch
- pitch is the psychophysical counterpart of freq
- 2 major theories of how vibration affects the basilar membrane and the encoding freq (pitch)

> which fibres are responding - Place Theory
- specific groups of hair cells on basilar membrane activate specific set of nerve fibers (guitar string)

> How fibres are firing- freq theory at Temporal code
- rate or pattern of firing of nerve impulses
Code of Pitch: Place Theory
- Suggests that diff freq disturb diff regions of bas. membrane (ie, diff cells)
- Helmhotz proposed "resonance" theory
> bas mem. wider at apex
> suggested fibers in bas mem resonated at diff freq at diff locations along its length (long fibres at apex, short at base)

- Problem w/ resonance theory
> no fibers
> not under tension
Psychophysical Tuning Curves
>experimental procedure
- First, a low level test tone is presented
- Then, masking tones are presented w/ freq above & below test tone
- Measures are taken to determine level of each masking tone needed to eliminate perception of test tone
- Assumption is that masking tones must be causing activity at same locations as test tone for masking to occur - common neural mechanism

> Resulting tuning curves show that the test tone is affected by a narrow range of masking tones

> Psychophysical tuning curves show the same pattern as neural tuning curves which reveals a close connection between perception and the firing of auditory fibers.
Evidence for Place Coding

- Stimulation Deafness Experiments
- expose animals to high amplitude tones of a particular freq & examine which hair cells are damaged
- Low freq causes damage to cells near the end
> damage to relatively broad area

- High freq causes damage to cells near stapes
> damage to relatively narrow area
Frequency Theory (Temporal Code)
- Proposes the basilar membrane vibrates in synchrony w/ the pressure changes (at the same freq as the sound)
- Freq representation based on a match between the freq in incoming sound waves and the firing rate of auditory nerve fibers
- Similar to a diaphragm in a speaker
> vibrates as a unit
> results in impulses in the auditory nerve at same freq as sound itself

- Two Problems:
> bas membrane not like a diaphragm
> nerve fibres can't fire above 1000 Hz
Problems for theories of the encoding of Pitch
- loudness
- changes in pitch perception in noise
- missing fundamental
Loudness & pitch
- as the intensity of tone changes, so does the pitch
- low freq tones - high intensity sounds appear lower in pitch
- high freq tones - high intensity tones appear higher in pitch
shifts of pitch in noise
- tones heard in background noise - pitch appears to be shifted in direction opposite to the pitch of the noise
> tone in lower freq noise appears high in pitch
> tone in noise higher in freq appears lower in pitch
The missing fundamental
- most natural sounds have a fundamental freq and harmonics, which are always some multiple of the fundamental
- the defining characteristic of the sound is the fundamental
- the missing fundamental phenomenon occurs when only the higher harmonics of a tone are presented
- In this situation, the fundamental is still heard
- Poses problems for a simple place theory
Pitch encoding
- Is pitch encoded according to place or freq theory?
- likely a combination of both of these theories accounts for our perception of pitch
- but there is also evidence for a role of auditory cortex in the perception of pitch as well
Loudness Perception (amplitude representation)
Firing rate = freq x amplitude
- a similar "principle of univariance" in colour vision

Dynamic Range
- the range of amplitudes that can be heard and discriminated when applied to an individual nerve fiber, the range of amplitudes over which the firing rate of the fiber changes
Hearing Impairment
- decrease in a persons ability to detect or discriminate sounds compared to a healthy young adult
Tinnitus
- persistent perception of sound in one's ears, ringing or buzzing.
Myths about Deafness
- deaf people can't hear anything
- hearing aids "cure" deafness
- deaf people can't talk
- deaf people are less intelligent than hearing people
- all deaf people can read lips
- deafness (or hearing impairment) is not just a simple reduction in auditory sensitivity
- its not just having permanent ear plugs
Audiometer
- instrument that tests peoples hearing
- presents pure tones w/ known freq and amplitude to right or left of ear
- Audiologis uses staircase method to estimate persons absolute threshold for each 6-8 frequencies, ranging from 250-8,000 Hz

- need to take into acct freq & intensity
- represents amt of hearing loss compared to normal (audiometric zero)
Audiogram
Graphical depiction of auditory sensitivity to specific freq, compared to sensitivity of a standard listener. Used to characterize possible hearing loss
- can reveal the kind of hearing loss present: degree of hearing loss in dB

Hearing is measure under diff conditions
- air or bone conduction
air conduction:
> a sound goes through mid-ear

bone conduction:
> sound bypasses mid ear

- Masked or unmasked
> masking ensures that only one ear can hear the test tone
Forms of Hearing Loss

-2 types
> Conductive hearing impairment
- outer or mid ear
- usually overall reduction in sensitivity

> Sensory/neural loss
- inner ear or auditory neurons
- may show loss over entire range of audible freq or selective loss
Conductive Hearing Impairment

Mechanical impediment
> common causes:
- blockage in external ear
- otitis media (infection)
- otosclerosis
- punctured eardrum

Diagnosis
- similar bone and air thresholds
- typical high freq loss

Treatment:
- advanced stage, replacing stapes
- cochlear implants

- not considered profound (loss of 90dB or >) Sound can till be conducted to cochlea via vibrations in bones
Sensori-Neural Loss
Hearing impairment caused by damage to cochlea, auditory nerve, or auditory areas or pathways of the brain

Common Causes:
- age, noise, ototoxic (drugs)

Diagnosis:
- similar bone and air thresholds
- loss may be profound

Treatment:
- hearing aids
- cochlear implants
Prebycusis
- age related hearing impairment
- life long exposure to noise
- long exposure to: drugs, chemicals, diabetes, CV disease, head trauam
- greater in men than women