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122 Cards in this Set

  • Front
  • Back
Meristem
Tissue in which cells are actively dividing.
Types of Apical Meristem types:
Initials - One remains meristematic.
Derivatives - becomes a new body cell.
Apical meristems bring about...
primary growth - growth in elongation.
Secondary growth
Growth in diameter.
3 overlapping processes for maturation of plant.
Growth - division and enlargement.
Morphogenesis - development of a shape or a form.
Differentiation - How cells differ based on location and gene expression.
Primary tissues
Make up the primary body.
Tissue
group of cells working together to perform a certain function.
Tissues are separated into three tissue systems:
Dermal
Vascular
Ground
Permanent tissue
Tissues, after being produced by meristematic tissue, that has function.

All tissues of the three tissue systems are permanent tissues.
Permanent tissues are either...
2 things.
Simple - all cells performing function are same type.
EX: Ground tissues - Parenchyma, Schlerenchyma, collenchyma
Complex - two or more cell types working together to perform function.
EX: Vascular, Dermal
Simple Tissues:
Composed of three types of cells.
Schlerenchyma, Collenchyma, Parenchyma
Parenchyma:
1. Characteristics
2. Location.
3. Function
4. Types
1. Living, Primary Cell Wall
2. Throughout
3. Metabolism, Storage, Conduction (from cell to cell)
4. Transfer
Aerenchyma
Chlorenchyma

MOST COMMON CELL TYPE IN PLANT BODY
Chollenchyma:
1. Characteristics
2. Location
3. Function
4. Types
1. Elongated, Unevenly thickened primary cell wall, occur in groups
2. Beneath epidermis
Ex: Celery strings
Leaf petioles
3. Support
4. Angular
Sclerenchyma
1. Types
2. Characteristics
3. Location
4. Function
1. Sclereids, fibers
2. Shape varies, secondary cell walls
3. Throughout (leaves, stems, fruits, seed coats, etc)
4. Protection and support
Types of Sclereids
Brachysclereids
Astrosclereids
Columnar
Osteosclereids
Fibers
1. Characteristics
2. Location
3. Function
1. Elongated, .2mm to .5 mm
Thick secondary cell wall.
Dead.
2. Xylem and Phloem
Throughout (ex. - many leaves of monocots)
3. Supports
Complex tissues
Vascular (xylem and phloem)
Dermal (epidermis)
Xylem: the principal conducting cells.
Tracheary elements, composed of two types:
Tracheids and vessel elements.
Tracheids: Found in which type of plant?

Have pits? perforations?
Seedless vascular plants, gymnosperms, primitive angiosperms.

Have pits, do not have perforations.
Vessels: Found in which type of plant?

Have pits? Perforation plates?
Angiosperms

Have both pits and perforation plates.
Types of plates
(4)
Scalariform - perforation plate
Simple - No perforation bars
Angular - go at an angle?
Truncate - most advanced, cut straight across
Which is more advanced, fewer or more bars on perforation plate?
Fewer bars.
Phloem:
Types of cells
Parenchyma
Fibers
Sieve elements, composed of a) Sieve cells and b) sieve-tube elements
Types of Sieve elements are their corresponding plant types.
Sieve cells - Seedless vascular plants and gymnosperms

Sieve Tube elements - Angiosperms
Sieve Cells
Characteristics
Primary cell wall
Living
Lack a nucleus
Pore Size - Sieve Area (group of pores)
Albuminous Cell
Specialized parenchyma cell associated with each sieve cell.

Regulates loading/unloading of sugars into sieve cell.

Albuminous cell and associated sieve cells are derived from different mother cells.
Callose
Polysaccharide composed of glucans.
Once cell is injured, releases glucans to block openings.

Protective mechanism against insects, continuous loss of sugar.
Sieve tube elements
Characteristics
Common in angiosperms.
Primary cell wall.
Living.
Lack a nucleus.
Pore size - Sieve plate
Compound vs. Simple
(simple is more advanced)
Companion cell
Associated with each sieve tube elements, they are parenchyma cells, helps with loading/unloading.

Come from the same parent cell.
P-protein
Once called "slime."

Forms a slime plug when the plant is cut. Prevents other plant nutrients from exiting.
Epidermis
Characteristics
Cover primary plant body
Parenchyma cells
Outer wall thickest (the one exposed to the environment)
Lack chloroplasts (most)
Living at maturity (typically)
Epidermis
Function
Protection
Prevents water loss/Absorption
Support
Specialized Epidermal features
Cuticle (cutin)
Prevents water loss
Mechanical barrier
Synthesized by epidermal cells
Lacking or ill formed in roots
Epicuticular waxes
Platelets
Rods
Granules
Hollow
Types of epidermal cells:
Ordinary
Very ordinary?
Types of epidermal tissues
Guard Cells
Specialized epidermal cells that occur in pairs.
Poorly developed or lacking cuticle.
Can change shape.
Contain openings called stomae.
How gases are exchanged.
Transpiration - loss of water vapor by the plant
Transpiration
Loss of water VAPOR by the plant
Types of epidermal tissues:
Subsidiary cells
Specialized epidermal guard cells that occur around guard cells
Types of epidermal tissues:
Bulliform cells
enlarged epidermal cells that can collapse or expand. Allows leaf to fold or roll up.
Types of epidermal tissues:
Trichome
Provide protection, reduces water loss by reducing amount of air that can blow across leaf.
Shoot
Consists of the stem and its leaves
Primary growth of shoot
Apical Meristem (most)
Intercalary meristems (few - grasses)
Secondary Growth of shoot
Vascular cambium, cork cambium
Apical Meristem
1. AKA
2. Characteristics
3. Function
1. Terminal bud
2. Covered by bud scales - protect apical meristem
3. Responsible for primary growth. Produce leaf primordia. Produce axillary buds.
Apical Meristem Histology:
Cyto-Histological Concept
Majumdar 1942
Gymnosperms
Tunica Corpus Concept
Angiosperms
Tunica
Outer most layer of cells
Divide anticlinally
Responsible for surface growth
Most angiosperms have 3 layers. L1, L2, L3
Corpus
Spherical body of cells beneath tunica.
Divide in various planes
Add bulk to developing stem
Two zones of Corpus
Peripheral/Marginal Zone
-outer layer of corpus
-lots of cells divisions

Central mother zone
-Bulk of corpus
-Few cell divisions
Peripheral Meristem
From l1, l2, l3 and peripheral zone of corpus.
Pith or Rib Meristem
Area just beneath central mother cell zone
Develops from peripheral zone.
Tissue System Development:
Dermal
L1
Ground
L1, L2, L3
Pith Meristem
Tissue System Development:
Vascular
L1, L2, L3
Peripheral Meristem
Pith Meristem
Elongation of stem
In active apical meristem, leaf and bud primordia develop very fast so it is difficult to distinguish between nodes and internodes.
Modes of Internode elongation
1. Uniform division and elongation throughout
2. Wave of divisions from base upward
3. Divisions restricted to base of internodes (intercalary meristem)
Increase in stem thickness during primary growth due to:
Periclinal cell division
Cell enlargement
Xylem (enarch)
From inside toward outside.
Protoxylem.
Metaxylem
-cell diameter larger
Phloem (exarch)
Outside to inside.
Protophloem
Metaphloem
Procambium
Vascular cambium
Xylem or Phloem
Protoxylem
Matures while stem is still elongating.
Often destroyed after metaxylem matures.
Wall thickenings are annular and helical.
Protophloem
First to mature.
Look like elongated parenchyma cell without nuclei.
Protoxylem
Development
Annular
Helical
Metaxylem
Development
Scalariform
Pitted
Continuous Vascular Bundle

1. Types of plants, example
2. Regions w/ type of cell
1.Magnoliids and Primitive Eudicots
Ex. tilia

2. a) Interfascicular region
-parenchyma cells
b) Vascular cambium (single layer)
-parenchyma cells
c) Cortex
-Parenchyma and Collenchyma
d) Pith
-Parenchyma
e) Ducts or Canals
-Mucilage (slimy carbohydrates)
Ring of separated vascular bundles
1. In what kind of plants
2. What two types of pith rays
1.Most Eudicots
2. Open, Closed
Open Pith Ray
Single layer of vascular cambium
Intrafascicular Vascular Cambim - Vascular Tissue
Interfascicular Vascular Cambium - Ground Tissue
Occurs in plants that have secondary growth.
Ex: Sambucus and Medicago
Closed Pith Ray
No vascular cambium
Vascular bundles surrounded by a sheath of sclerenchyma cells.
Occurs in plants that lack secondary growth.
Ex: Ranunculus and Helianthus.
Scattered Vascular Bundles
Some Eudicots
All Monocots
V.B. closed
Lack secondary growth
EX: Zea
Nodal Anatomy:
1. Stem bundle
2. Leaf Trace
3. Branch Trace
1. vascular bundle of stem
2. vascular tissue that extends from stem bundle out into the leaf
3. vascular tissue that extends from stem bundle out into lateral bud
1. Leaf Gap
2. Branch Gap
3. Sympodium
1. break in vascular tissue above leaf trace.
2. break in vascular tissue above branch trace
3. stem bundle and associated leaf trace
Which plants have secondary growth?

Which do not have secondary growth?
Gymnosperms
Magnoliids
Some Eudicots

Monocots
Most plants that have secondary growth are classified as
Perennials - plants that live for more than two years.
What happens to primary tissues when secondary tissue growth occurs?
Primary tissue is destroyed.
Secondary growth:
Lateral Meristems
(2)
1. -Vascular Cambium
-Secondary xylem
-secondary phloem
-vascular rays
2. Cork cambium
Periderm - replaces epidermis when destroyed.
Intrafascicular

Interfascicular
From vascular tissue

From ground tissue
Planes of cell division
1. Periclinal
2. Anticlinal
1. Parallel to surface
2. perpendicular to surface
Fusiform Initials
Vertically elongated
Gives rise to vascular tissues:
1. Vessels or tracheids
2. Sieve cell or sieve tube elements
3. Parenchyma
4. Fibers
Ray Initials
Horizontally elongated
Gives rise to vascular rays
Parenchyma
Secondary xylem rays
Secondary phloem rays
Functions of Vascular Rays:
1. Lateral Transport. Sucrose symplastically (across plasma membrane to living part of cell to next cell)
Water apoplastically (along/between cells, does not cross plasma membrane)

2. Storage tissue
-Starch
-Lipids
Vascular Ray Classification
*Seen in tangential section (face view)
1. Uniseriate
2. Biseriate
3. Multiseriate
1. One cell layer wide
2. Two cell layers wide
3. 3 or more cell layers wide
Secondary Phloem
Phloem transport, young/old
Functional phloem - younger secondary phloem involved in transport
Non functional phloem - no longer involved in transport
Secondary Xylem
Wood of tree
Types of Section Cuts
1. Transverse section

(longitudinal)
2. radial (radius of tree)
3. tangential (face view of radial section)
Growth rings
1. What do they do
2. Spring wood
3. Summer wood
4. Fall wood
1. Tell age of tree in years, esp in temperate zones
2. Springwood - large vessels, lots of transport to make leaves, take up more water
3. Summer wood - leaves have been produced cell diameter decreases
4. Fall wood - cell diameter continues to decrease, becomes dormant
Ring Porous wood
Vessles in spring wood very large
Diffuse porous -
vessels tend to be uniform in diameter throughout growth ring (though there are more present during spring, summer)
Non-stoned and stoned wood:
Non-stoned - overlaps of end walls, cells end at different places, stronger

Stoned wood: cells tend to occur in groups, ending at same places. Weak points.
Heartwood
Often darker portion in center. Older secondary xylem, no longer involved in transport.
Sapwood
lighter in color.
Younger secondary xylem, actively involved in transport
How sapwood changes into heartwood
Changes occur before death in living cells.
1. food reserves removed
2. oils, gums, resins formed
3. Tyloses form (in many species)
-balloon like outgrowth from ray parynchyma cells into vessels.
-growth through pits of vessel
-Consist of plysaccharides and pectin
-Line inner walls of heartwood
Softwood
A longitudinal tracheid. Thicker, elongated.
3-5 mm long.
Writing paper and brown bags.
Hardwood
B. vessel (early wood) - short, wide
D. Vessel (late wood) - short, narrow
C. Fiber (1-2 mm long - very narrow, elongated
E. Tracheid - a little bit longer and narrower than vessels
-kodak paper, toilet paper, napkins, kleenex
Knots
Where a lateral branch has been covered by growth rings
Types:
Tight
Loose
Suberin
fatty material that lines wall of phellum cells
Cork Cambium
Origin
In stems, comes from one of the outer layers of cortex.
In roots, from pericycle.
Phellum
Cork. Results from division of Cork Cambium, to the outside. (dead, suberin)
Phelloderm
Formed from division of Cork Cambium, to the inside. Living at maturity.
Periderm
Replaces the epidermis as protective covering. Consists of phellum, phellogen, phelloderm.
Lenticel
opening in periderm for gas exchange.

Formed by cells growing faster than other cells, creates a break in the periderm.
What does bark consist of?
Periderm
Secondary phloem
Leaves
Primary plant organ produced by a primary meristem
Leaves occur on current year's growth only
True leaves are Megaphylls
Leaf Development
Clonal analysis
Marker cells
Chimeral Meristems - Apical meristem injected with marker cells
Leaves initiate from a group of cell sin the peripheral meristem. From l1, l2, l3 layers.
These cells (5-100 are founder cells) elongate.
Grass leaf development
Marginal meristem encircles the stem. Blade wraps around stem. Sheath-part wraps around
1.Leaf buttress
2. Leaf primordium
3. marginal meristems
1. the bump, when a new leaf starts forming.
2. Start to see some symmetry
3. Marginal meristems - cause widening of the leaf. Arise from founder cells.
Vascular tissue development
Major veins develop upward and outward as the midrib develops.
Minor veins develop from tip towards the base.
Tip of leaf fully developed first.

Primary, Secondary, Tertiary, Quaternary growth - ascending in order, getting smaller and finer.
Phyllotaxy
1. Alternate
2. Opposite
3. Whorled
How leaves are arranged on a stem.
1. One leaf per node.
-Helical (45 degrees)- evolved so that each leaf can absorb maximum sunlight
-Distichous - (180 degrees) - opposite sides.
2. Decussate - two leaves per node
3. Whorled - 3 or more leaves per node
AKA Verticillate
What determines phyllotaxis?
1.Classical model.
2. New Model
1.Hormone inhibitor prevents formation of leaf primordia.

2. Auxin (Indole acetic acid) induces new primordia. These primordia serve as a sink and drain IAA from surrounding cells.
Leaf Parts
1. Petiole
2. Lamina
3. Stipule
4. Sessile
5. Sheath
6. Ligule
7. Abaxial
8. Adaxial

Leaflet anatomy
9. Petiolule
10. Rachis
1. Stalk of leaf
2. Blade of leaf
3. Small leaflike structures found at edge of leaf
4. Leaf with no petiole
5.When base of leaf wraps around stem (grasses, etc)
6. Thin membrane, holds sheath onto stem
7. Underside of leaf
8. Upper surface of leaf
9. Stalk of a leaflet.
10. Area of central axis between leaflets
Simple Vs Compound leaves - Lamina
Simple - lamina is one piece.

Compound - Lamina is broken down into small units aka leaflets.
Types of Compound Leaves:
Palmately
Leaflets all originate from common point like palm of hand.
Types of compound leaves:
Pinnately compound
Like pinnae of a feather. Axis with radiating leaflets.
Types of Compound Leaves
Bi-pinnately compound
(A form of pinnately compound)
Secondary leaflet
Secondary Petiolule
Rachilla - Secondary Rachis
Leaf Anatomy:
Tissue Systems
Dermal - Epidermis
Ground - Mesophyll, Bundle Sheath
Vascular - Xylem, Phloem
Epidermis:
1. Parenchyma cell characteristics
2. Functions
1. Living at maturity (normally)
Xerophyes (secondary c.w. and dead)
Outer wall thickest
Lack chloroplasts

2. Prevent water loss
Protection (mechanical and chemical)
Support
Specialized epidermal features
Cuticle (made of cutin)
Epicuticular waxes
Trichomes
Bulliform cells
Guard cells
Guard cells
specialized epidermal cells that occur in pairs. Has opening called stoma. Stoma is only opening in epidermis. Allows gas exchange of co2 and O2. Allows for transpiration.
Remiform guard cells
Kidney shaped. Eudicots.
Ostidform guard cells
Bone shaped. Monocots.
Subsidiary cells
Morphologically distinct epidermal cells immediately surrounding guard cells.
Anomocytic cells
Subsidiary cells surrounding the guard cell pair are not morphologically distinct from other epidermal cells
Stomata location
1. Hypostomatic
2. Amphistomatic
3. Epistomatic
4. Astomatic
1. Guard cells on lower surface
2. guard cells occur on both upper and lower surface
3. Guard cells occur on upper surface. Leaves that float on surface of water.
4. No guard cells. For leaves that are submerged.
Guard cell position
1. Above
2. Equal
3. Sunken
4. Stomata crypt
1. hydrophytes
2. mesophytes
3. Xerophytes
4. sunken pit with hairs, guard cells tucked away in pit.