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48 Cards in this Set

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Due to the extreme environments plants live in, structures that could adapt evolved.

For protection, retention of water, entry restriction of organisms and toxic substances, and strengthening and supporting the plant.

Epidermis definition

Superficial layer of cells of all parts of the primary plant body, but not found in the root cap and not differentiated as such on apical meristem.

Epidermis origin

Outermost cell layer of apical meristem/tunica layer (if tunica-corpus present)




Roots: may be related developmentally to cortex or root cap

"Protoderm"

Undifferentiated epidermis, regardless of origin




Haberlandt's term: "primordial epidermis" or epidermis from separate initials

Retains epidermis?

Organs with little or no secondary growth.

Epidermis with one layer cells in thickness

Uniseriate

More than one layer cells in thickness; Due to what?; Example.

Multiseriate; Periclinal division/unsynchronized division layers; Velamen of orchid plants

First protective barrier between aerial surface of plant and environment + principal barrier to movement of water.

Cuticle




Note: superficial, but if in terms of cells, epidermis is superficial.

General structure of plant cuticle.

Epicuticular wax


Cuticle proper


Cuticular layer


Pectin


Cell wall

Factors that affect cuticular permeability.

Structure and amount of wax

2 lipid components of cuticle

1. Insoluble cutin - in the matrix


2. Soluble cutin - some at the surface; some in the matrix called cuticular wax

Cuticularization

Secretion of cutin to surface of cell wall

Cutinization

Deposition of cutin between cellulose microfibrils of outermost layers, where pectin and hemicellulose may also be present.

Distinct region formed by cuticularization; Presence of cellulose?

Cuticle proper; No cellulose

Distinct region formed by cutinization

Cuticular layer

Layer under cuticular layers is rich in?; Continuous with?

Pectin; middle lamella

Structural composition: epidermal cell proper

-least specialized, ground mass


-mostly parenchyma cells


-tabular cells


-deficient in chlorophyll (plastids with few grana)

Has morphologic variability @ epidermal cell proper

Gramineae (grasses)



(*) morphologic variability in cells due to?

Long cells




Short cells


a.) silica cells - contains silicon dioxide (smaller)


b.) cork cells - suberized walls (larger)





Bulliform cells: brief description

"bubble-like", thin-walled, highly vacuolated, water containing cells

Bulliform cells: walls

thin radial walls




outer walls as thick/thicker than those of adjacent ordinary epidermal cells




composed of cellulose and pectic substances




outer walls are cutinized + bear cuticle

Bulliform cells: expansion tissues

cause rapid expansion of leaf

Bulliform cells: motor cells

cause changes in turgor in opening/closing movement of leaf

Other specialized epidermal cells

Lithocysts

Guard cells


cells turgid? ; cells flaccid?

stoma open; stoma closed

Guard cells control diameter of stoma. How?

Changing shape thru osmosis. (note: constant length)




Taking in water = more turgid (due to orientation of cellulose microfibrils, cells buckle outward) = open




Losing water = more flaccid = close



Cause of change in turgor pressure that open and close stomata

Result primarily from reversible uptake and loss of K+ by guard cells


-uptake of K+ from adjacent epidermal cells to vacuole: osmotic flow of water due to decreased water potential


-loss of K+: osmotic loss of water

Stomatal rims or ledges

elevated extensions of cuticular membrane

Types of stomates (dicot)

Anomocytic (subsidiary cells = ordinary cells)


Anisocytic (3 SCs)


Diacytic (2 SCs, perpendicular)


Paracytic (2 SCs, parallel)

Types of stomates (monocot)

Type I (4-6 SCs; usually "boxed in")




Type II (4-6 SCs; rounded top+bottom and smaller than lateral SCs)




Type III (paracytic)




Type IV (anomocytic)

Guard cells: Gramineae and Cyperaceae (describe)

dumb-bell shaped; when turgid, expanded ends swell, pulling the elongated parts apart (open)

Stomata in some xerophytes

Located @ abaxial leaf surface. Usually found in stomatal crypts.

Stomata in some xerophytes: stomatal crypts

sunken areas of leaf surface to protect pores from dry winds

Stomata in some xerophytes: trichomes

"hair"; minimize transpiration by breaking up flow of air keeping crypt more humid than in the surrounding atmosphere

Sunken stomate

Subsidiary cells overarch guard cells (i Pinus i) or formed epidermis with thick cuticular wall (i Franklandia i)

Stomata positions

amphistomatic (both surfaces)


epistomatic (upper surface)


hypostomatic (more commonly on lower surface)

Stomatal development: stomata arise through?

Differential division of protodermal cell

Stomatal development: guard mother cell

-product of the division; immediate precursor of guard cells


-2 guard cells arise after 1-3 divisions


-occur @ same level as adj epidermal cells; rising or sinking of stomate occurs during maturation

Stomatal development: pores

-before pores separate, area where they form contains pectic enzymes


-formation is initiated enzymatically



stomatal development: separation of guard cells

occur thru starch hydrolysis

(*) Stomatal development: anisocytic

asymmetric division of EC => meristemoid + non-stomatal EC1 => 2nd division (meristemoid) => EC2 => 3rd division (M) => EC3 + Guard mother cell => division of GMC => 2 guard cells, no pore yet

Trichomes: brief description

-hair


-epidermal appendages


-may occur on all parts of plant; may persist or fall off early

Trichomes: functions

-reduces transpiration rate by increasing boundary layer (still air thru which water vapor must diffuse)


-defense against insects


-secretory may provide defense

Dendritic trichome

Branched along an axis

Glandular trichome @ stinging hair of i Urtica i

Pressure on unsilicified base forces liquid into wound




contain histamine, acethylcholine, serotonin (irritation maybe)

Common type of trichome

Scale or peltate hair: found in mint, shield-shaped cluster of cells

Secretion: Internal secretory structures

Idioblasts (tannin, calcium oxalate)




Lithocyst




Laticifers

Secretion: external secretory structures

Hyathodes-modified stomates where excess water is released thru guttation




*epithem-parenchyma tissue lacking chlorophyll between vein and external pore




glandular trichomes