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152 Cards in this Set

  • Front
  • Back

Clade

a group of organisms containing the last common ancestor and all of its descendants

Metazoa

a scientific word for "animals"

Protostome means "________________"

mouth first

Deuterostome means "_______________"

mouth second

2 sub-clades of the super-clade Protostomia

Lophotrochozoa and Ecdysozoa

Lophotrochozoa

lophophore feeding apparatus (ring of hollow tentacles) and/or a trochophore larvae, a type of motile larvae with a band of cilia around the middle

Ecdysozoa

animals that molt, shed part of their body, at some point in their development

Edward Wilson

"The Father of Biodiversity"


professor emeritus at Harvard


written many books

Geological time in the circular cladogram starts with what era in the middle? which is the outermost layer?

inside - Hadean Era


outisde - middle Cenozoic Era

describe the formation of the cladogram

Time moves outward beginning with the Hadean Era in the innermost layer and Cenozoic on the outermost layer. The time of origin is where the branches begin. The ovals represent phyla. The width of the ovals is relative to the amount of species in that phyla. Some phyla are broken up into smaller phyla

Hadean Era

4.6 billion years ago when the Earth formed. Ended around 3.8 billion years ago when the planet began to cool. Name in greek means "underworld."

Archaean Era

3.8 billion years ago to 2.5 billion years ago.


the first prokaryotes evolved (bacteria and archaea)


these bacteria and archaea make up 2/3 major domains of life

Prokaryote

organisms that lack a cell nucleus and any other membrane-bound organelles

Eukaryotes

organisms that contain a cell nucleus and first appeared in the Proterozoic Era

Proterozoic Era

first eukaryotes


2.5 billion - 543 million years ago


"base of the Cambrian Period"


Metazoans are first observed in the fossil record


radiation of animals

Cambrian period

600 million - 500 million years ago


every phyla that is extant today evolved during this period


Extant

alive today

3 domains of life

bacteria


archaea


eukaryotes

Opisthokonts

the clade that contains fungi, choanoflagellates and the animals (in the eukaryote domain of life)

Animals are _____________, ______________, _____________, are capable of __________ at some point in development, and have regulatory genes called _____________ genes.

multicellular, heterotrophic, lack a cell wall


movement


Hox genes

Hox genes

only found in Metazoan clade


-a group of genes that specify anterior-posterior axis and segment identity during early embryonic development


-ONLY animals have Hox genes

animals are multicellular - meaning?

differentiated cells for specialized functions but is not a defining feature of animals

Heterotroph

ingest and digest their food but not a defining characteristic of animals

Capable of movement at some stage of development

larval stage sponge moves but adult doesn't


not a defining feature of an animal

no cell wall

protists also lack a cell wall

What is biodiversity?

-biodiversity is the totality of genetic diversity, species diversity, ecosystem diversity in a region

Species Diversity

number of species in a region and their relative abundances

Species Richness

number of species in a region


(Earth has many millions of distinct biological species)

What percent of species richness do vertebrates make up?

about 5%

What percent of species richness do invertebrates make up?

about 95%

Pea-sized Pony

-worlds smallest seahorse


-named about 3 years ago


-just over 1cm

Barbados Thread Snake

under 10cm


worlds smallest snake

Phylum Arthropoda make up what percentage of metazoan species?

about 83%

How many Metazoan Phyla are there?

34

2 historic definitions of a phylum

-as a group of organisms with a certain degree of morphological or developmental similarity (phenetic definition)


-as group of organisms with a certain degree of relatedness (phylogentic definition)

Convergent Evolution

describes the acquisition of the same biological trait in unrelated lineages


-analogous structures or homoplasy

Divergent Evolution

opposite of convergent, related secies evolve different traits


-homologous structures

Stem Group Organisms

lack one or more features that are present at the base of the crown group t which they are attached


all members of the stem-group of a clade are extinct

Crown Group

contains the last common ancestor of all extant members and all of the ancestor's descendants

Stem Group diagram

-vertical line represents a clade of phylum


-thin lines are stem members


-thick lines are crown members


-red dots are first appearance of crown member


-red horizontal lines show stem members that evolved pre-Cambrian

Extremophiles

organisms that live in harsh environments such as hot springs, salt lakes



first Archaea were extremophiles

Stromatolites

are layered accretionary structures that form in shallow water


formed by trapping and binding of sedimentary grains by biofilms of microorganisms, especially cyanobacteria

Rangeomorphs

have branching "frond-like" elements. Each element is itself composed of many smaller elements

During the Cambrian Period, what was life like?

all life was aquatic, most life was relatively small, many animals had unusual body layouts

Sanctacaris

A large shrimp that grows to 9cm


An arthropod


A chelicerate

Chelicerate

-Invaded earth more than 400 million years ago


-arthropod group that includes spiders, scorpions, and horseshoe crabs.

Hallucegenia

close relative of a rare modern animal known as onychophoran

Onychophoran

-share certain characters with arthropods, but are lacking a hard exoskeleton or jointed legs


-not a diverse group, less than 100 species extant today

Pikaia

-thought to be a worm


-now thought to be a chordate due to rod running along back

Opabinia

-5 mushroom-shaped eyes, clawed proboscis, and unusual body


-evolved alongside arthropods, chordates and echinoderms but it's lineage went extinct


-might be closely related to arthropods (stem group)

4 broad habitat types

Marine


Freshwater


Terrestrial


Symbiotic or Parasitic

2 marine/freshwater habitats

benthic


pelagic

Benthic

Bottom

Pelagic

Top (water column)

2 Terrestrial habitats

Xeric (dry)


Moist

Parasitic habitat

Negative effect on organism that is habitat


e.g. Roundworm

Symbiotic Habitat

Positive effect on both organisms


e.g. Cleaner fish (Phyla Chordata)

2 Parasitic/Symbiotic Habitats

Endo-parasites


Ecto-parasites

In grid of 34 phyla, the colours mean what?

Green: 1-100 species


Blue: 100-1000 species


Yellow: 1000-10000 species


Orange: 10000-100000 species


Red: 100000+

What phyla have been sub-divided in the diagram?

Arthropoda (crustacea, chelicerata, and uniramia)


Chordata (urochordata and cephalachordata)

What habitats contain the most phyla?

Aquatic (marine and freshwater)


29/34 phyla

Of the 5 phyla that aren't found in aquatic habitats, how many are not symbiotic/parasitic?

1 (Onychophora)

How many phyla can be found in the marine habitat? How many can be found in the freshwater habitat?

29, 14

In the marine habitat, how many phyla can be found in the benthic habitat? pelagic?



In the freshwater habitat, how many phyla can be found in the benthic habitat? pelagic?

29, 14



14, 6

What phyla can be found in all three habitats?

Nematoda, Arthropoda, and Chordata


(3/34)

How many phyla have diversified to more than 100 000 species?

2 (Arthropoda and Mollusca)

What are the 5 key transitions?

Tissue


Symmetry


Body Cavity


Development


Segmentation

Cladogram

a diagram which shows ancestral relations between organisms an represents the evolutionary tree of life

First branch of metazoan cladogram

Phylum Placozoa from other metazoans

What is the second branch of the metazoan cladogram?

Porifera from all other organisms. (Poriferans don't undergo gastrulation)

Gastrulation

is a phase early in the development of most animal embryos, during which the morphology of the embryo is reorganized to form the germ layers: ectoderm and endoderm in diploblasts and ectoderm, endoderm and mesoderm in triploblasts.

What is the third split?

Radiata (Cnidaria and Ctenophora, radial symmetry) and Bilateria (bilateral symmetry)

What id the evolutionary difference between protostomes and deuterostomes?

blastopore becomes the mouth = protostome


blastopore becomes the anus = deuterostome

Protostomes branch into how many super-clades? What are they called?

2


the Lophotrochozoa (protostomes with a trochophore larvae or lophophore feeding apparatus) and the Ecdysozoa (molting animals)

Grades of Biological Organization

Cellular


Cell-Tissue


Tissue


Tissue-Organ


Organ

Cellular Biological Organization

see aggregation of cells that are functionally differentiated

Cell-Tissue Organization

see a cellular organization where we have ensemble of cells that are functionally differentiated

Tissue Grade of Organization

ensemble of cells that carry out a specific function

Tissue-Organ Grade of Organization

aggregation of tissues


first seen in Acoelomorpha

Organ Grade of Organization

organs working together to perform a function


highest grade of organization

Symmetry

the balanced distribution of duplicate body parts or shapes

Bilateral Symmetry

have a front and a back end, as well as an upside and downside

Assymmetry

no axis of symmetry

Radial Symmetry

topside (aboral) and downside (oral) but no front or back

Bi-radial Symmetry

has both bilateral and radial axes of symmetry

Pentaradial Symmetry

5-sided symmetry (starfish)

Acoelomate

an animal that lacks a coelom

Pseudocoelomate

2 body cavities formed but the pseudocoelum is not fully surrounded by a mesoderm so not a true cavity

Ceolomate Body Plan

mesoderm splits into a true body cavity called the ceolom (developed in triploblasts but were lost in several lineages)

Triploblasts

have a body made of 3 cellular layers, the exterior ectoderm and the interior endoderm and a middle mesoderm

What is another word for segmentation?

metamerism

Segmentation

is the repetition of a similar body segments along longitudinal axis of body

What is each segment called?

metamere or somite

What is true metamerism?

involves repetition of a structure with common function derived from both ectoderm and mesoderm

What is Serial Repetition?

if the repetition only involves the ectoderm such as integument (skin) of some playhelminths such as the repeating nature of the proglottids in tapeworms (pink body sections) or the repeating nature of the integument of some of the nematodes.

How many times has metamerism independently evolved? In what?

3


annelids, arthropods, and chordates

Porifera

Sponges and Coral


"pore-bearers"


aquatic (mostly marine), benthic


9000 species named (about 100 in freshwater)


no pelagic sponges


sessile, live on substrate


some are ectosymbionts or commensalists

Placozoa

Marine


only 1 species named

What are the 5 key transitions of Porifera?

Tissue: cell-tissue


Symmetry: assymmetrical


Body Cavity: no body cavity


Development: no germ layers (protostome)


Segmentation: no

What are the 5 key transitions of Placozoa?

Tissue: cell-tissue


Symmetry: assymmetrical


Body Cavity: no body cavity


Development: no germ layers (protostome)


Segmentation: no

Sponge Body is assembled of what?

Gelatinous Matrix

Common characteristic of all sponges

a body made up of a complex system of canals and pores which are open to the surrounding water

Hexactinellida

glass sponge


body made up of siliceous spicules with 6 rays

Calcarea

strictly marine


mostly in shallow tropical waters


body made up of calcium carbonate


3 body grades: asconoid, synconoid and leuconoid

Demospongiae

largest class of sponge


secrete a form of collagen called spongin and secretes siliceous spicules


"bath sponge"


90% of all sponges

Homoscleromorpha

class of marine sponge made of 2 families


84 species in this class

Poriferans have a unique canal system called what? What does it do?

Aquiferous system


-system of canals and chambers connected to the surrounding environment by multiple pores


-made of small incurrent pores (ostia) and 1 or more excurrent pores (oscula) and a spacious spongocoel.

The sponge body has 2 primary layers of cells. What are they called?

Pinacoderm (outer layer)


Choanoderm (inner layer)


(between the two layers is a mesohyl)

Mesohyl

the gelatinous matrix and is the connective tissue of the sponges - in it are found the ameoboid cells and the skeletal elements

Asconoids

simplest body type; water enters through the ostia then goes to large internal chambers called the spongocoel (lined with cells called collar cells or choanocytes which are the main filter feeding apparatus of the sponge); asconoids are typically small - all under 10 cm and are tube shaped and attached to substrate; all are in the class Calcarea.

Synconoids

thicker than asconoids; unlike the asconoid the layer where the spongocoel is found is folded into radial canals; the canals are lined with choanocytes; water enters radial canals through tiny openings called prosopyles; the water empties into spongocoel; all synconoids are in the class Calcarea

Leuconoids

most complex; most efficient at pumping water; increased complexity of aquiferous system allows the sponge to not just pump more water but capture more food thus it is usually larger; diverse body type; have flagellated chambers; no spongocoel; demospongiae are all this type

The sperm and eggs of sponges are produced by what?

choanocytes

4 types of Placozoan cells

1) monociliated dorsal cells


2) ventral epithelia cells


3) ventral gland cells


4) syncytail fiber cells

Choanocytes

make up the principle "pump" and "filter" of the system, driving water through the sponge, trapping and phagocytizing suspended bacteria and other particulate food

Archaeocytes

are amoeboid cells that can transform into all other cell types (totipotent cell)

What are the 5 Key Transitions of Cnidaria?

Tissue: tissue


Symmetry: radial symmetry


Body Cavity: no body cavity


Development: diploblasts; protostrome


Segmentation: no

What are the 5 Key Transitions of Ctenophora?

Tissue: tissue


Symmetry: bi-radial symmetry


Body Cavity: no body cavity


Development: diploblasts; protostrome


Segmentation: no

What habitats are Cnidaria and Ctenophora found in?

Solely aquatic (primarily marine); benthic or pelagic

What are the body plans of Cnidarians?

benthic polyp form



pelagic medusae form

The polyp body form resembles a flower and stalk. Why?

The stalk is a cylindrical column arising from the petal disk.



At the oral end is the manubrium: and elevation with the mouth at its summit situated in the centre of the oral disk

What shape is a medusa?

an umbrella or bell

Cnidarians have an outer __________ and an inner ________________.

epidermis; gastrodermis

What are the 5 classes of Cnidarians and the distinguishing features if each? What features do all Cnidarians share?

All share: radial body form, cnidocytes, planula larvae, and a mouth surrounded by solid tentacles. Anthozoa (gut with septal filaments, siphonoglyph, anthozoan pharynx, hexaradial and octaradial symmetry), Staurozoa (creeping planula without cilia), Scyphozoa (rhopalium; polyp reduced or lost), Cubozoa (velarium), Hydrozoa (polyp lost, velum in medusae, medusae produced by lateral budding and enntocodon)

The external anatomy of the "classic" ctenophore is characterized by what?

a sac-like body resembling a peeled grape

General function of nervous systems:

to collect and process information about the surrounding environment, then to produce appropriate actions

General nervous system trend:

a progression from diffuse nerve nets to highly centralized systems with differentiated sensory organs

The first nerves recognizable at the cellular level are found in the ______________. Nerves allow cells to transfer information from one cell to another. This is done in ___________ -- either through direct ______________ (electrical synapses) or via ______________ transmitters (chemical synapses).

Cnidarians


synapses


transmission


chemical

Cnidocyte

endodermal interstitial cells



occur through the epidermis and are most abundant on the tentacles

Cnidocytes bear 2 important organelles. What are they?

Nemacysts (a capsule with an inverted coiled tube and operculum)



Cnidocil (trigger for the ejection of the nematocyst)

Ctenophores have what instead of cnidocytes?

collecytes (burst open when prey comes in contact with the tentacles, sticky threads released from each of the colloblasts will then capture the food)

Statocysts

gravity-sensing structures



highly sensitive structures lined with sensory hairs, which transmit a signal when the hairs are bent.

What 3 features do Ctenophores share with Cnidaria?

1) diploblast organization with extensive, mostly non-cellular middle layer (mesoglea)


2) possess only a mouth (no true anus; but some species in both phyla have anal pores)


3) gastrovascular cavity with radiating canals

5 major differences between Ctenophores and Cnidaria

1) Asexual reproduction not well-developed


2) all are hermaphroditic but external cross fertilization is typical


3) use ctenes for movement rather than sheets of epitheliomuscular cells or myocytes


4) lack cnidocytes (instead have collecytes)


5) swim with their mouth forward

Platyhelminthes lives in what habitat?

Moist Terrestrial, Marine, Freshwater

Acoelomorpha lives in what habitat?

Marine (benthic and pelagic)

Which phyla (acoelomorpha or platyhelminthes) have many classes that are parasitic?

Platyhelminthes (ecto- and endo-parasites)

What are the 5 key transitions of Acoelomorpha?

Tissue: tissue-organ


Symmetry: bilateral symmetry


Body Cavity: acoelomate


Development: triploblasts; protostome


Segmentation: no

What are the 5 key transitions of Platyhelminthes?

Tissue: tissue-organ


Symmetry: bilateral symmetry


Body Cavity: acoelomate


Development: triploblasts; protostome


Segmentation: no

What does A-P axis stand for?

Anterior-Posterior Axis

Cephalization:

is an evolutionary trend, whereby nervous tissue, over many generations, becomes concentrated toward one end of an organism


forms a head region

Cephalization is intrinsically connected with a change in __________?

symmetry

Why is the combination of cephalization and bilateral symmetry important in animals?

it allows the sensory organs to face the direction of movement

What are the photoreceptors called in acoels?

"ocelli"


cup-shaped pigment cells that can sense the direction of light

Tubellaria

one of the traditional sub-divisions of the phylum Platyheminthes, and include all of the sub-groups that are not exclusively parasitic



most forms reproduce sexually and with few exceptions

Planaria

non-parasitic flatworms with ability to regenerate body parts

Neodermata contain what 3 parasitic groups?

Cestoda, Trematoda, Monogenea

Cestodes are known as ___________?

tapeworms

Trematoda are known as ____________?

flukes

In what 3 ways do the 3 classes of Neodermata differ from free-living turbellaria?

-lack sense organs


-epidermis lacks cilia


-have a weird syncytial epithelium with microvilli

Scolex

The worm's head attaches to the intestine of the definitive host.

Proglottids

successive segments that compose the body