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137 Cards in this Set

  • Front
  • Back
What are the Two general categories of the classical theories of hearing?
place, frequency
The place theory was postulated by ___________
Helmholtz
Classical theory that postulated that structures acted like finely tuned resonators. Implied that basilar membrane is composed of a series of tuned segments that resonate to particular frequencies. The part of the membrane corresponding to the incoming stimulus was the part that vibrated.
Place Theory
What were the three main problems with the Place theory?
BM tension, missing fundamentals, tuning
In order for the Place theory to work and to account for the sharp tuning of the inner ear, it would be necessary for those finely tuned resonators to be under differing amounts of ______________.
tension
(Place Theory) Is the basilar membrane under any tension?
No
(Place Theory) The missing fundamental was thought to be a result of distortion that was caused by the nonlinearity of middle ear. Is the middle ear actual nonlinear?
No
(Place Theory) What the true cause of the missing fundamental?
Ear uses both frequency pattern and temporal pattern to determine pitch. The ear uses the fundamental frequency of a combination of tones as well as the period of the fundamental frequency in order to determine what frequency we are hearing.
(Place theory) In order for the basilar membrane to respond to specific frequencies at specific locations, the basilar membrane would have to have __________ tuning, _________ damping, and ________ ringing, which would in turn cause echoes. This disproves the place theory because we do not hear echoes in average speech.
narrow, low, long
The theory that states that the cochlea is NOT frequency specific but responds to all frequencies all along the basilar membrane. After responding to the frequencies, the hair cells transmit frequencies to auditory nerve for analysis
Frequency (Telephone) Theory
What are the problems with the frequency theory
There is a limit to the number of nerve fiber discharges per second so the frequency theory cannot amount for frequencies over 1KHZ. If the frequency theory cannot amount for frequencies over 1 KHz, it also cannot account for why high frequency hearing loss is associated with certain parts of the basilar membrane
What kind of response does the action potential have?
all or nothing
The refactory period which states that once the action potential fires it can't fire again for another 1 ms.
absolute refractory period
The refractory period after the absolute refractory period that requires a greater amount of stimulus in order for the nerve to fire again.
relative refractory period
Combination of the Place/Frequency (Telephone) Theories that suggests that one neuron doesn’t have to carry all of the information alone, but that groups of fibers work together.
Volley Theory
an observed phenomenon (in support of the volley principle) where neurons fire in synchrony with the phase of a stimulus. This works for frequencies of up to _______________ Hz
phase locking, 4000-5000
No individual neuron could fire at each peak, but a bunch of phase-locked neurons working together can produce a burst of activity at each peak, and so the firing frequency of a collection of neurons can indeed mimic the __________________ of the stimulus
frequency
Below 300-400 Hz ________________ likely predominates
temporal coding
Phase locking works for frequencies up to ___________________ Hz
4000-5000
Above 4000-5000 Hz ________________ takes predominates
the place theory
Between 400 and 4000 __________ takes predominance
A combination of volley & place theories
sound transmitted to inner ear via_____________
ossicular chain
The sound transmitted to the inner ear by the ossicular chain causes stapes to rock in and out of __________, causing fluid within cochlea to move as a wave. This is known as the _________________ theory.
oval window, traveling wave
Wave is propagated along basilar membrane from _______ to ________, which causes bm to be displaced. Basilar membrane becomes displaced in a wave like motion=___________. The basilar membrane also becomes displaced from _________ to __________
base, apex
traveling wave
base, apex
Basilar membrane displacement _____________ in amplitude until a _____________ displacement is reached and then _____________abruptly
increases, maximum, decreases
The greater the stimulus level (intensity level of stimulus) the greater the _____________
displacement
Displacement peak occurs near apex for _______ freq and near base for ________ freqs.
low, high
Displacement peak occurs near _________ for high freq and near ________ for low freqs.
base for high freq, apex for low freqs
Displacement is ALWAYS from ________ to _________, regardless of frequency
base, apex
For very ______ freqs, displacement occurs along nearly the entire basilar membrane, but peaks near apex
low
The input frequency determines not only the ______________ that the traveling wave moves, but the rate of basilar membrane __________.
distance, vibration
The basal end can vibrate to ______________ freqs and the apical end can vibrate to __________freqs
basal= low and high freqs
apical= only low freqs
High freq stimulation results in large displacements at ________
base
Low freqs cause small displacements at __________first & large displacement at __________
basal end, apex
The traveling wave is how the cochlea performs ______________________ analysis
frequency
Each hair cell contains sensory receptors or____________
stereocilia
Each hair cell contains sensory receptors or stereocilia.
Cilia from _______ are embedded in the tectorial membrane; whereas cilia from ______ are not
OHCs, IHCs
When the basilar membrane is deflected, the stereocilia are bent and the hair cells are activated. Bending of the stereocilia ___________ the modiulus activates the hair cells and __________ firing rate of aud. neurons
away from, increases
Bending __________ the modiolus inhibits the hair cells and ____________ firing rate
towards, decreases
What are the two main functions of the cochlea?
1. provide frequency analysis on incoming signals
2. convert mechanical energy to neuroelectric energy via the HC and synapse with auditory neurons
Generated by nerve fiber (acoustic 8th cn); Recorded from the nerve fiber itself or sites near nerve; All-or-none potential that appears as a spike
Action potential
Single nerve fiber responses are recorded by inserting a __________________ into a nerve fiber
microelectrode
the time it takes the action potential of the auditory nerve fibers to reach maximum (amp) potential & return to resting state
spike
The number of times the nerve fires (discharges) per second is called the ____________
firing rate
Nerve fibers require no stimulation and can fire on their own. This is known as the _______________ rate.
spontaneous
A nerve fiber is _____________ when its firing rate increases above its spontaneous rate by some percentage (e.g. 20%).
activated
The minimum intensity level needed for a nerve fiber to become activated and increase its firing rate above its spontaneous rate is the nerve fiber's ____________.
threshold
The lowest intensity that gives rise to a criterion percentage increase in rate at which the fiber is firing above its spontaneous rate is the nerve fiber's _________________
threshold
Nerve fibers can respond to more than one frequency depending on the ___________ level
stimulus
The frequency at which the nerve fiber responds best is called the ___________________
characteristic frequency
Fibers with High characteristic frequencies are found in hair cells in the
base of cochlea
Fibers with low characteristic frequencies are found in hair cells in the
apex of cochlea
Increasing stimulus intensity increases _______________ until a maximum rate is reached (usually _________ dB above the spontaneous rate.). In this way the nerve fiber can encode _____________
spike rate, 30-40, intensity
For single nerve fibers, as the stimulus intensity increases, the ______________ increases, but the ___________ does not
spike rate, amplitude of the spike
Chart that reflects the most frequent time interval between each pair of neural discharges/indicates that the most frequent interspike interval corresponds to the period of the waveform
interval histogram
Interval histograms prove that fibers help to encode the _________ of the waveform.
period
occur as the result of a large group of neurons firing
whole nerve action potential/ compound action potentials
WN action potentials can be recorded by positioning surface ____________ on the head and other body parts.
electrodes
Whole nerve action potentials can be recorded from locations __________ to the nerve
distant
Are WN responses an all-or-none event?
No
What kind of response are WN or CAP action potentials?
graded
With WN/CAPh, the more nerve fibers firing, the greater the _____________
amplitude
Brief stimulus durations such as clicks are used to increase the ___________________ of nerve fiber responses and thus increase the _____________ of nerve fiber responses
synchronization, amplitude
After receiving the stimuli, the traveling wave takes ________ ms to travel the full length of the cochlea from the base to the apex.
2-4 ms
__________________ fibers respond synchronously to stimuli and thus provide the largest contribution to the whole-nerve action potential.
Basal high frequency
The response of the WN/CAP are measured in terms of ____________ and ______________
latency, amplitude
The interval b/t stimulus onset and the onset of the neural response
latency
As the stimulus level increases, the latency ___________ and the amplitude ___________
decreases, increases
The study of the relationship between the sound stimulus and the response it produces in the subject
psychoacoustics
What are the two methods of measurement used by psychoacousticians?
Discriminating procedures
Scaling techniques
Procedure that measures where a listener has to make subjective response to presence of a signal
Discriminating procedures
Procedure that analyzes the smallest difference that would allow the listener to discriminate between 2 stimulus conditions
Discriminating procedures
What is the end result of the discriminating procedure?
threshold
The threshold that describes the lowest stimulus level that we can present a signal to you and you can detect it presence from no stimulus being present. You should be able to detect this 50% of the time
Absolute threshold
The absolute threshold detects the ____________ of a signal.
presence
The threshold that indicates the smallest change in a stimulus that can be detected by the listener
Difference Threshold
Threshold that is used to notice/detect the smallest difference in frequency or intensity between two tones a certain % of the time.
Difference Threshold.
Techniques that measure behavioral responses to intensities or frequencies.
Scaling techniques
Scaling techniques attempt to measure sensation directly by measuring what two parameters?
loudness and pitch
What are the three measures of discrimination procedures that are used to measure absolute and difference thresholds?
Method of limits
Method of adjustments
Method of constant stimuli
Test in which there are several trials. Will present the signal at a # of diff intensity levels. Trials alternate between ascending and descending. The stimulus adjustment is not continuous. Experimenter makes the adjustment. The bigger the adjustment, the bigger the chance for error in the threshold.
Method of Limits
Similar to method of limits except for the listener/participant adjusts the stimulus level. The stimulus adjustment is continuous.
Method of Adjustment
The experimenter presents the stimulus level at varying intensities. But there is a constant stimuli=When the experiment is finished, the stimulus level has been presented at each stimulus level the same number of times. Ex: The experimenter will present stimulus at 2, 4, 6, and 8 db 3 times each.
Method of Constant Stimuli
Modified versions of the three types of discrimination procedures which help to prevent biased results as a result of factors other than the magnitude of sensation (such as by biasing the subject or incorrectly administering procedures)
Adaptive Procedures
Test that quantifies a subjective attribute of sound such as loudness or pitch by allowing the experimenter to determine set of stimulus parameters that all yield the same subjective sensation.
Matching Procedures
Behavioral responses elicited in a sound field (both ears exposed to the sound)
Minimal audible field
Behavioral responses obtained under earphones, sound stimuli presented monaurally (one ear at a time)
Minimum audible pressure
Is hearing equally sensitive across frequencies?
No
More SPL is necessary to elicit responses at the ______________ of the audibility range and again at the _______________ of the range
lower end, higher end
Hearing sensitivity in the human ear ranges from ____ to _________ Hz
20, 2000
The missing dB occurs because the This is because the responses are better in the___________ than they are in the ___________.
sound field, headphones
The psychological correlate to intensity is known as ____________
loudness
What is the missing 6 db?
Refers to the difference in thresholds obtained in the sound field vs. under the earphones
The first efforts to establish equal loudness levels occurred in or around ___________
1927
The earliest measures to assess relative loudness was developed by _________and _________ (1933). The curves are sometimes called the _______________ curves or _____________________
Fletcher and Munson
Fletcher-Munson curves
equal loudness contours
Curves that tell us how loud the intensity of a tone of one freq must be to = the intensity of a tone of another freq. at 1000 Hz at a particular level
Equal Loudness Contours
The curve created when presented tone at 40 dB and took another tone and adjusted the tones until the person listening said that they were equal in tone. Thus these tones became equal.
40 phon curve
The unit of level for equal loudness
Phon
(Loudness) Required ________ energy at low levels to equal intensity of high frequency levels
more
Loudness grows faster for _______ frequencies.
low
Ask the listener to make comparisons between a standard stimulus and a reference stimulus by assigning numbers according to how the listener perceives the stimulus
Scaling Procedures
Unit of measurement for loudness for a 1000 Hz tone at 40db SPL
Sone
The reference for one sone is ______ dB SPL
40
One Sone= _______ Phons
40
Stephens Power law says that the the sensation of Loudness (L) increases as a power (e) of the _______________ L=kI ^e
stimulus intensity
(Stephen's Power Law) If loudness grew at the same rate as stimulus increased, the power of the stimulus intensity would be ________
1.0
If loudness is growing at a rate slower than the intensity change, than the power function/slope will be _______ than one.
less
If loudness is growing at a faster rate than the intensity change, then the slope will be _________ than one.
greater
For Signals less than 30 phons, slope was steeper because loudness grows faster for ______ frequencies.
low
Signals less than 30 phons, slope was ___________ because loudness grows faster for low frequencies.
steeper
For every spl increase of about 10 decibels, we are sensing a __________ of loudness.
doubling
If a sound was twice as loud as a 1000 Hz tone at 40 dB SPL it is _____ sones
2
If a sound was half as loud as a 1000 Hz tone at 40 dB SPL it is _____ sones
.5
The processing of sound by two ears
Binaural Hearing
Difference between the time it takes signal to reach left ear vs time it takes to reach the right ear
Interaural time difference
For Lower freqs below 1500 hz, the interaural time difference will encode in an interaural _______________ difference
phase
The ear that is stimulated _____ signals the direction of the sound
first
The magniute of the interval time difference increases as the location of sound source changes from ___________ to __________
straight ahead (0 degrees) to straight out to the side (90 or 270 degrees)
When the sound originates directly in front of the listener, the length of the pathway to the ears is _________
equal
When sound originates directly in front of the listener does an interaural time difference exist in the arrival of sound?
No
At the extreme right or left (90 or 270 degrees), the difference between the length of the path to the near ear and far ear is greatest. This equates to a ________ interaural time difference.
maximum
An interaural time difference of ___ ms occurs for all frequencies.
.5
For a pure tone with a frequency of 1000 Hz that completes one cycle in 1 ms, the signal to the far ear would start _____ cycle after the signal to the near ear. The two signals would have a _______ degree phase difference between the two ears.
.5, 180
A pure tone of 500 Hz with a 2 ms period would only be delayed _____ of the period. This would result in a _______ degrees interaural phase difference
1/4, 90
Although interaural time difference is _______ for all frequencies, interaural phase difference is _____ among frequencies.
equal, varying
Interaural ___________ Difference results from a sound shadow.
Intensity
Sound shadow is produced when wavelength of sound is ________ relative to dimensions of the object
small
Sound shadow created by head ___________ with frequencies above 500 Hz.
increases
At 6000 Hz for 90 degrees and 270 degrees, the interaural intensity difference results in a ______ dB difference.
20
The intensity of the near ear is _______ dB greater than intensity at far ear.
20
At 500 Hz interaural intensity difference is less than _____ dB
4
At 0 degrees, the interaural intensity difference is ___ dB for all frequencies
0
As you increase frequency, intensity difference is _________
greater
For low freqs intraaural ________ difference is dominant cue
time
For higher frequencies intraaural ____________ difference is predominant cue
intensity
Duplex theory of sound localization at horizontal plane: the two primary cues for localization of sound in space are what?
interaural time difference and interaural intensity differences
Both cues may be used over a wide range of freqs by listeners to identify the location of the sound source, but at low freqs interaural _____ difference predominates and at high freqs interaural _______ differences predeominates
time, intensity